Landscape Ecology Lab
Betts Forest Landscape Ecology Lab

Home | Research | Recent Press | Applications | Teaching | Publications | People | Lab News | Positions


Matt Betts on Google Scholar

For pdfs of the publications below, see Researchgate

Students from the Betts lab are highlighted in red, postdocs in green.


Strauss, S.H., Jones, K.N., Lu, H., Petit, J.D., Klocko, A.L., Betts, M.G., Brosi, B.J., Fletcher, R.J. and Needham, M.D. In Press. Genetically engineered reproductive modification in trees: potential biodiversity impacts and social dimensions. New Phytologist.

Ripple, W. Abernethy, K., Betts, M.G., Chapron, G., Dirzo, R., Galetti, M. Levi, T., Lindsey, P.A., Macdonald, W., Machovina, P., Newsome, T.M., Peres, C., Wallach, A.D., Wolf, C., Young, H. 2016. Bushmeat hunting and extinction risk to the world's mammals. Royal Society Open Science 3: 160498
Featured in: The Huffington Post,, Science, Mongabay, The Guardian

Root, H.T., Verschuyl, J., Hammond, P., Stokely, T., Scherr, M. and Betts, M.G. In Press. Plant diversity enhances moth diversity in an intensive forest management experiment. Ecological Applications.

Comfort, E.J., Clark, D.A., Anthony, R.G., Bailey, J. and Betts, M.G. In Press. Quantifying edges as gradients at multiple scales improves habitat selection models for northern spotted owl. Landscape Ecology

Kroll, A.J., Giovanini, J., Verschuyl, J. and Betts, M.G. In Press. Assembly dynamics of a forest bird community depend on disturbance intensity and foraging guild. Journal of Applied Ecology

Frey, S.K.J., Hadley, A.S. and Betts, M.G. 2016. Microclimate predicts within-season distribution dynamics of mountain forest birds. Diversity and Distributions: 1-16.
Featured in: NW Climate Science Center

Hadley, A.S. and Betts, M.G. 2016. Refocusing habitat fragmentation research using lessons from the last decade. Current Landscape Ecology Reports: 1: 1-12

Frey, S.J.K., Hadley, A.S., Johnson, S. L., Schulze, M., Jones, J. A. and Betts, M. G. 2016. Spatial models reveal the microclimatic buffering capacity of old-growth forests. Science Advances 2, e1501392.
Featured in:,, Tech Times, Gazette-Times, Science Daily

Briggs, F., Fern, X., Betts, M.G. and Raich, R. 2016. Multi-instance multi-label class discovery: a computational approach for assessing bird biodiversity. Proceedings of the AAAI Conference on Artificial Intelligence (AAAI-16).

Kormann, U., Scherber, C., Tscharntke, T., Klein, N., Larbig, Valente, J.M., Hadley, A.S. and Betts, M.G. 2016. Corridors restore animal-mediated pollination in fragmented tropical forest landscapes. Proceedings of the Royal Society Ser. B. 283: 1823.
Featured in: Science Daily,, Conservation Magazine,

Root, H. and Betts M.G. 2016. Managing moist temperate forests for bioenergy and biodiversity. Journal of Forestry 114(1): 66-74.


Givot, R., O'Connell, K., Hadley, A.S. and Betts, M.G. 2015. Hummingbird citizen science. The Science Teacher NOVEMBER(08): 25-33.

Schmidt, K.A., Johansson, J. and Betts, M.G. 2015. Information-mediated Allee effects in breeding habitat selection. American Naturalist 186: E162-E171

Moriarty K.M., Epps C.W., Betts M.G., Hance D.J., Bailey J.D., Zielinski W.J. 2015. Experimental evidence that simplified forest structure interacts with snow cover to influence functional connectivity for Pacific martens. Landscape Ecology 30: 1865-1877

Hutchinson, R.A., Valente, J.V., Emerson, S.C., Betts, M.G., and Dietterich, T.G., 2015. Penalized Likelihood methods improve parameter estimates in occupancy models. Methods in Ecology and Evolution.

Betts, M.G., Hadley, A.S. and Kress, W.J. 2015. Pollinator recognition by a keystone tropical plant. Proceedings of the National Academy of Sciences of the USA 112:3433–3438. doi: 10.1073/pnas.1419522112
Featured in New York Times, Nature, AAAS Science Update, CBC Quirks and Quarks, Scientific American

Betts, M.G., Gutzwiller, K.J., Smith, M.J., Robinson, W.D. and Hadley, A.S. 2015. Improving inferences about functional connectivity from animal translocations experiments. Landscape Ecology 30: 585-593. DOI 10.1007/s10980-015-0156-x


Bennett, V.J., Betts, M.G. and Smith, W. 2014. Influence of thermal conditions on habitat use by a rare spring emerging butterfly Euphydryas editha taylori. Journal of Applied Entomology 138: 623–634.

Volpe, N.L., Hadley, A.S., Robinson, W.D. and Betts. M.G. 2014. Functional connectivity experiments reflect routine movement behavior of a tropical hummingbird species. Ecological Applications 24: 2122–2131.
See article summary in the Bulletin of the Ecological Society of America pp. 161-164: website

Gutiérrez Illán, J., Thomas, C.D., Jones, J.A. Wong, W.K., Shirley, S.M. and Betts, M.G. 2014. Precipitation and winter temperature predict long-term range-scale abundance changes in Western North American birds. Global Change Biology 20(11): 3351–3364.
Featured in Science, Science World Report, Nature World News

Betts, M.G., Fahrig, L., Hadley, A.S., Halstead, K.E., Robinson, W.D. Bowman, J., Wiens, J.A., and Lindenmayer, D.B. 2014. A species-centered approach for uncovering generalities in organism responses to habitat loss and fragmentation. Ecography 37(6):517-527.

Hadley, A.S., Frey, S., Robinson, W.D., Kress, J. and Betts, M.G. 2014. Tropical forest fragmentation limits pollination of a keystone understory herb. Ecology 95(8): 2202-2212.

Pfeifer (and 54 authors including Betts, M.G.) 2014. BIOFRAG – A new database for analysing BIOdiversity responses to forest FRAGmentation. Ecology and Evolution 4(9): 1524-1537. Supporting Information

Rivers, J.W., Liebl, A.L., Martin, L.B., and Betts, M.G. 2014. Corticosterone in territorial male Swainson's Thrushes varies in relation to forest age but not vegetation cover. Journal of Ornithology 155(2): 539-548.


Briggs, F., et al. 2013. The 9th annual MLSP competition: New methods for acoustic classification of multiple simultaneous bird species in a noisy environment. IEEE International Workshop on Machine Learning for Signal Processing, MLSP

Bennett, V.J., Pack, S.M.; Smith, W.P. and Betts, M.G. 2013. Sex-biased dispersal in a rare butterfly and the implications for its conservation. Journal of Insect Conservation 17(5): 949-958.

Smith, M.J., Forbes, G.J., and Betts, M.G. 2013. Landscape configuration influences gap-crossing decisions of northern flying squirrel (Glaucomys sabrinus). Biological Conservation 168: 176-183.

Betts, M.G., Verschuyl, J., Giovanini, J., Stokely, T., and Kroll, A.J. 2013. Initial experimental effects of intensive forest management on avian abundance. Forest Ecology and Management 310: 1036-1044.

Shirley, S.M., Yang, Z., Hutchinson, R.A., Alexander, J.D., McGarigal, K., and Betts, M.G. 2013. Species distribution modelling for the people: unclassified landsat TM imagery predicts bird occurrence at fine resolutions. Diversity and Distributions 19: 651-872.

Jenkins, S.R., Betts, M.G., Huso, M.M., and Hagar, J.C. 2013. Habitat selection by juvenile Swainson’s thrushes (Catharus ustulatus) in headwater riparian areas, Northwestern Oregon, USA. Forest Ecology and Management 305: 88-95.

Yegorova, S., Betts, M.G., Hagar, J., and Puettmann, K.J. 2013. Bird-vegetation associations in thinned and unthinned young Douglas-fir forests 10 years after thinning. Forest Ecology and Management 15: 1057-1070.

Cahall, R.E., Hayes, J.P., and Betts, M.G. 2013. Will they come? Long-term response by forest birds to experimental thinning supports the ‘‘Field of Dreams’’ hypothesis. Forest Ecology and Management 304: 137-149.


Jones, J., Kroll, A.J., Giovanini, J., Duke, S.D., Ellis, T.M., and Betts, M.G. 2012. Avian species richness in relation to forest management practices in early seral tree plantations. PLoS One 7(8): e43290.

Kroll, A.J., Duke, S.D., Hane, M.E., Johnson, J.R., Rochelle, M., Betts, M.G., and Arnett, E.B. 2012. Landscape composition influences avian colonization of experimentally created snags. Biological Conservation 152: 145-151.

Rivers, J.W., Liebl, A.L., Martin, L.B., and Betts, M.G. 2012. Baseline corticosterone is positively related to juvenile survival in a migrant passerine bird. Functional Ecology 26(5): 1127-1134.
Featured in Science Daily

Ellis, T.M., Kroll, A.J. and Betts, M.G., 2012. Early seral hardwood vegetation increases adult and fledgling bird abundance in Douglas-fir plantations of the Oregon Coast Range. Canadian Journal of Forest Research 4: 918-933.

Briggs, F., Lakshminarayanan, B., Neal, L., Fern, X, Raich, R., Frey, S.K., Hadley, A.S. and Betts, M.G. 2012. target="_blank"Acoustic classification of multiple simultaneous bird species: a multi-instance multi-label approach. Journal of the Acoustical Society of America 131: 4640-4650.
Featured in

Swatantran, A. Dubayah, R. Goetz, S. Hofton, Betts, M.G., Sun, M., Simard, M. and Holmes, R.T. 2012. Mapping migratory bird prevalence using remote sensing data fusion. PLoS One 7: e28922

Hadley, A.S. and Betts, M.G. 2012. The effects of landscape fragmentation on pollination dynamics: absence of evidence not evidence of absence. Biological Reviews 87(3): 526-544


Smith, M.J., Forbes, G.J. and Betts, M.G. 2011. Evidence of multiple annual litters in Glaucomys sabrinus (Northern Flying Squirrel). Northeastern Naturalist 18(3): 386-389.

Smith, M.J., Betts, M.G., Forbes, G.J., Bourgeois, M.C., Flemming S.P. and Kehler, D.G. 2011. Independent effects of connectivity predict homing success by northern flying squirrel in a forest mosaic. Landscape Ecology 26: 709-721.

Jones, J., Kroll, A.J., Giovanini, J., Duke, S.D., and Betts, M.G. 2011. Estimating occupancy thresholds with imperfect species detection. Ecology 92: 2299-2309.

Zitske, B.P., Betts, M.G. and Diamond, A.W. 2011. Negative Effects of Habitat Loss on Survival of Migrant Warblers in a Forest Mosaic. Conservation Biology 25: 993-1001.

Ellis, T. and Betts, M.G. 2011. Bird abundance and diversity across a hardwood gradient within early seral plantation forest. Forest Ecology and Management 261: 1372-1381.

Bennett, V.J., Betts, M.G., Smith, W.P. 2011. Mate guarding behaviour in the male Taylor’s Checkerspot butterfly. Journal of Insect Behaviour 23: 165-250.

Rivers, J.W., Martin, L.B., Liebl, A.L. and Betts, M.G. 2011. Parental alarm calls of the white-crowned sparrow fail to stimulate corticosterone production of their nest-bound offspring. Ethology 117: 374-384.


Betts, M.G., Hagar, J., Rivers, J. Alexander, J. McGarigal, K. and McComb, B.C. 2010. Thresholds in forest bird occurrence as a function of the amount of early-seral broadleaf forest at landscape scales. Ecological Applications 20: 2116-2130.

Goetz , S.J., Steinberg, D., Betts, M.G., Holmes, R.T., Doran, P.J. Dubayah, R. and Hofton, M. 2010. Lidar remote sensing variables predict breeding habitat of a Neotropical migrant bird. Ecology 96: 1569-1576.
Featured in NSF-Science, Science Daily

Ahlering, M.A., Arlt, D., Betts, M.G., Fletcher, Jr., R.J. Nocera, J.J., and Ward, M.P. 2010. Research needs and recommendations for the use of conspecific-attraction methods in the conservation of migratory songbirds. Condor 112(2): 252-264.

Betts, M.G., Nocera, J.J, and Hadley, A.S. 2010. Settlement in novel habitats induced by social information may disrupt community structure. Condor 112(2): 265-273.

Nocera, J.J. and Betts, M.G. 2010. The role of social information in avian habitat selection. Condor 112(2): 222-224.

Wuest, L, and Betts, M.G. 2010. Quantitative tracking of the vegetative integrity and distinctness of forested ecological communities: A case study of plantation impacts. Canadian Journal of Forest Research 40: 330-346.


Rota, C., Fletcher, R., Dorazio, R. and Betts, M.G. 2009. Occupancy estimation and the closure assumption. Journal of Applied Ecology 46: 1173-1181.

Hadley, A.S. and Betts, M.G. 2009. Tropical deforestation alters hummingbird movement patterns. Biology Letters 5: 207-210
Featured in Science, Science Daily, The Oregonian

Ritchie, L., Betts, M.G., Forbes, G.J., and Vernes, K. 2009. Effects of landscape composition and configuration on northern flying squirrels in a forest mosaic. Forest Ecology and Management 257: 1920-1929.

Wilson, D., Stoddard, M.A., Betts, M.G., and Puettmann, K.J. 2009. Bayesian small-area models for assessing wildlife conservation risk in patchy populations. Conservation Biology 23: 982-91.

Betts, M.G., and M.-A. Villard. 2009. Landscape thresholds in species occurrence as quantitative targets for forest management: generality in space and time? In Setting Conservation Targets for Managed Forest Landscapes. Villard, M.-A. (Ed.) Cambridge University Press.

Betts, M.G., Ganio, L., Huso, M., Som, N., Bowman, J., Huettman, F. and Wintle, B. 2009. Comment on "Methods to account for spatial autocorrelation in the analysis of species distributional data: a review". Ecography 32: 374-378.


Betts, M.G., Hadley, A.S., Rodenhouse, N.L. and Nocera, J.J. 2008. Social information trumps vegetation structure in breeding site selection by a migrant songbird. Proceedings of the Royal Society of London-Ser. B. 275: 2257-2263
Featured in Conservation Magazine, Washington Post, Science Daily

Betts, M.G., Rodenhouse, N.L., Sillett, T.S. Doran, P.D. & Holmes, R.T. 2008. Dynamic occupancy models reveal within-breeding season movement up a habitat quality gradient by a migrant songbird. Ecography 31(5): 592-600.


Betts, M.G., Mitchell, D., Diamond, A.W., Bêty, J. 2007. Uneven rates of landscape change as a source of bias in roadside wildlife surveys. Journal of Wildlife Management 71: 2266-2273.

Betts, M.G., Forbes, G.J. and Diamond, A.W. 2007. Thresholds in songbird occurrence in relation to landscape structure. Conservation Biology 21: 1046-1058.

Linke, J., Betts, M.G., Lavigne, M. and Franklin, S.E. 2007. Introduction: Structure, function and change in forest landscapes. In Forest Disturbance and Spatial Pattern: Remote Sensing and GIS Approaches. Wulder, M. and Franklin, S.E. (Eds.) Taylor and Francis Group Publishing, Pp. 1-29.


Betts, M.G., Forbes, G.J., Diamond, A.W., and Taylor, P.D. 2006. Independent effects of habitat amount and fragmentation on songbirds in a forest mosaic. Ecological Applications 16: 1076-1089.

Betts, M.G., Diamond, A.W., Forbes, G.J., Villard, M.-A. and Gunn, J. 2006. The importance of spatial autocorrelation, extent and resolution in predicting forest bird occurrence. Ecological Modelling 191: 197-224.

Betts, M.G., Zitske, B., Hadley, A.S., and Diamond, A.W. 2006. Migrant forest songbirds undertake breeding dispersal following timber harvest. Northeastern Naturalist 13: 531-536.


Young, L., Betts, M.G., and Diamond, A.W. 2005. Do Blackburnian Warblers select mixed forest? The importance of spatial resolution in defining habitat. Forest Ecology and Management 214: 358-372.

Warren, T., Betts, M.G., Diamond, A.W. and Forbes, G.J. 2005. The influence of local habitat and landscape composition on cavity-nesting birds in a forested mosaic. Forest Ecology and Management 214: 331-343.

Betts, M.G., Hadley, A.S. and Doran, P.J. 2005. Avian mobbing response is restricted by territory boundaries: experimental evidence from two species of forest warblers. Ethology 111: 821-835.

Doran, P.D., Gulezian, P.Z. and Betts, M.G. 2005. A test of the mobbing playback method as a means to estimate bird reproductive success. Journal of Field Ornithology 76: 227-233.

Betts, M.G., Diamond, A.W., Forbes, G.J., Frego, K., Loo, J., Matson, B., Roberts, M., Villard, M.-A., Wissink, R., and Wuest, L. 2005. Plantations and biodiversity: a comment on the debate in New Brunswick. Forestry Chronicle 81: 265-269.

Betts, M.G., Simon, N.P.P. and Nocera, J. 2005. Point count data summary metrics differentially predict reproductive activity in forest and grassland birds. Journal of Ornithology 146: 151-159.


Betts, M.G., Franklin, S.E., and Taylor, R.G. 2003. Interpretation of landscape pattern and habitat change for local indicator species using satellite imagery and geographic information system data in New Brunswick, Canada. Canadian Journal of Forest Research 33: 1821-1831.


Betts, M.G., Knox, J. and Forbes, G.J. 2002. A landscape ecological approach to private woodlot planning in New Brunswick. Natural Areas Journal 22: 311-317.

Betts, M.G., and Loo, J. 2002. A comparison of pre-European settlement forest characterization methodologies. Forestry Chronicle 18: 123-129.