1. 2001. Common insects and diseases of interior Douglas-fir. British Columbia Ministry of Forests SIL471. 8 p.

Keywords:      tree/stand protection

                        tree/stand health

Abstract: This field guide provides information on the different pests and diseases of the interior Douglas fir (Pseudotsuga menziesii) in British Columbia which include: defoliating insects (Douglas fir tussock moth, spruce budworm and rusty tussock moth); dwarf mistletoe; foliar diseases (e.g., caused by the Cooley spruce adelgid); bark beetles; root diseases (Armillaria root disease, blackstain root disease, laminated root rot and blackstain root disease); wood decay (caused by bracket or conk fungi); and various abiotic problems (sunscald, drought or frost). A guideline to control infestations of these given pests is also included.

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2.
Adams, T., T. Anekonda and C. Lomas. 1999. Annual Report 1998-99, Pacific Northwest Tree Improvement Research Cooperative. 33 p.

Keywords:      genetic tree improvement

                        tree/stand protection

                        tree/stand health

                        growth

                        tree physiology

Abstract: Summaries are given of research projects on improvement of Douglas fir [Pseudotsuga menziesii] in the Pacific Northwest: seedling drought physiology; genetics of dark respiration and its relationship with drought hardiness; response of saplings to drought, as measured by growth ring variables; use of microsatellite marker loci to identify pollen contamination in seed orchards; and evaluation of miniaturized seed orchard designs.

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3.
Aitken, S.N. and W.T. Adams. 1996. Genetics of fall and winter cold hardiness of coastal Douglas-fir in Oregon. Canadian-Journal-of-Forest-Research 26(10): 1828-1837.

Keywords:      genetic tree improvement

                        tree/stand protection

                        tree/stand health

                        tree physiology

                        genetic relationships

Abstract: Genetic variation in autumn cold hardiness was studied in two western Oregon breeding populations of coastal Douglas fir (Pseudotsuga menziesii var. menziesii), one on the west slope of the Cascade Mountains and the other in the Coastal Range. On six sampling dates (September, October and November 1992 and January, September and October 1993), shoot cuttings from 40 open-pollinated families in each of two progeny test sites for each breeding zone were subject to artificial freezing at two test temperatures. Damage in each shoot was recorded as visible injury to needle, stem and bud tissue separately. Considerable family variation was found for cold injury scores in all tissues in early to mid autumn, but differences were often smaller or nonsignificant in late autumn and midwinter. Individual heritability estimates for needle cold injury were low (<0.40) and generally decreased in late autumn and midwinter. Family rankings for autumn cold hardiness, however, are expected to be relatively consistent over sites and years, although needles appear to display more family-by-site interaction than stems or buds. Genetic correlations between tissues in cold injury varied considerably and were sometimes weak, indicating that the evaluation of a single tissue is probably not adequate for assessing overall cold hardiness of genotypes. Autumn and winter cold hardiness seem to be largely under separate genetic control since genetic correlations between hardiness at these two stages were weak. This study confirms earlier results in Washington breeding populations and shows that coastal Douglas fir families can be effectively ranked for autumn cold hardiness by conducting artificial freeze tests on cut shoots in mid-autumn (October) and scoring damage to stems and at least one other tissue.

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4.
Aitken, S.N. and W.T. Adams. 1997. Spring cold hardiness under strong genetic control in Oregon populations of Pseudotsuga menziesii var. menziesii. Canadian-Journal-of-Forest-Research 27(11): 1773-1780.

Keywords:      genetic tree improvement

                        tree/stand protection

                        tree/stand health

                        tree phenology

                        tree physiology

                        genetic relationships

Abstract: Genetic variation in spring cold hardiness of shoots prior to bud break was studied in two Oregon breeding populations of Pseudotsuga menziesii var. menziesii, one on the west slope of the Cascade Mountains and the other in the Coast Range. In March and April 1993, and April 1994, shoot cuttings from 40 open-pollinated families in each of two progeny test sites in each breeding zone were subjected to artificial freezing. Visible cold damage to needle, stem, and bud tissues was recorded. Date of bud burst (all sites), and injury resulting from a 1992 natural frost event (one site), were also recorded. Spring cold injury varied widely among families. Individual heritabilities for spring cold injury scores averaged 0.76 in the Coastal zone and 0.42 in the Cascade zone. Genetic correlations among tissues, sites, sampling dates, and years, and between April cold injury and date of bud burst were high, in most cases over 0.80. Correlations were also strong between natural frost damage in 1992 and artificial cold injury scores in 1993. Artificial freeze testing stem tissues of cut shoots sampled in April from a single test site should effectively rank families in this region for spring cold hardiness.

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5.
Aitken, S.N., W.T. Adams, N. Schermann and L.H. Fuchigami. 1996. Family variation for fall cold hardiness in two Washington populations of coastal Douglas-fir (Pseudotsuga menziesii var. menziesii (Mirb.) Franco). Forest-Ecology-and-Management 80(1/3): 187-195.

Keywords:      genetic tree improvement

                        tree/stand protection

                        tree/stand health

                        tree physiology

                        tree phenology

                        genetic relationships

Abstract: In order to assess the genetics of autumn (fall) cold hardiness in coastal Douglas fir (Pseudotsuga menziesii var. menziesii), shoot cuttings were collected in October from saplings (9-year-old trees) of open-pollinated families in two progeny tests in each of two breeding zones in Washington, one in the Coast range (80 families) and one on the west slope of the Cascade Mountains (89 families). Samples from over 5500 trees were subjected to artificial freezing and visually evaluated for needle, stem and bud tissue injury. The extent to which cold injury is genetically related to tree height and shoot phenology (timing of bud burst and bud set) was also evaluated. Significant family variation was found for all cold hardiness traits; however, individual heritability estimates were relatively low (ranging from 0.09 to 0.22). Significant family-by-test site interaction was detected for needle injury in the Cascade breeding zone, but not in the coastal zone. Genetic correlations (rA) among needle, stem and bud tissues for cold damage were weak (0.16<less or =>rA<less or =>0.58) indicating that genes controlling autumn cold hardening are somewhat different for different tissues. Timing of bud burst and bud set were only weakly correlated with cold injury (rA<less or =>0.49). Thus, bud phenology is a poor predictor of autumn cold hardiness in this species. There was no consistent relationship between tree height and cold injury in the coastal zone. In the Cascade zone, taller trees appeared to be more susceptible to cold injury, but the association was weak (mean rA=0.38, range 0.20-0.72).

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6.
Alvarez, I.F. and R.G. Linderman. 1983. Effects of ethylene and fungicide dips during cold storage on root regeneration and survival of western conifers and their mycorrhizal fungi. Canadian-Journal-of-Forest-Research 13(5): 962-971.

Keywords:      nursery operations

                        tree/stand health           

                        mycorrhizal response

Abstract: Survival and growth of Pseudotsuga menziesii, Pinus ponderosa and Abies concolor seedlings, and survival of mycorrhizal fungi on their roots were assessed after cold storage with or without 5 p.p.m. ethylene in combination with 4 root treatments: washed, dipped in Truban [etridiazole] or Benlate sol. or not treated. Ethylene treatment resulted in increased survival, apical bud burst, and new root formation in the greenhouse if roots had not been washed or dipped in fungicide. None of the gas storage or root treatments greatly affected seedling survival in the field. Root washing decreased seedling vigour, especially in fir. None of the root treatments or gas storage conditions affected root fungal populations; bacterial and actinomycete populations appeared to be affected and the response varied according to host species. Pisolithus tinctorius, which formed mycorrhizae with 10-20% of the short roots of the seedlings, did not survive cold storage. Thelephora spp. and an ectendomycorrhizal fungus both survived cold storage and rapidly colonized roots newly formed on seedlings planted after cold storage.

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7.
Alvarez, I.F. and J.M. Trappe. 1983a. Dusting roots of Abies concolor and other conifers with Pisolithus tinctorius spores at outplanting time proves ineffective. Canadian-Journal-of-Forest-Research 13(5): 1021-1023.

Keywords:      planting operations

                        site preparation

                        mechanical preparation

                        growth

                        tree/stand health

                        mycorrhizal response

Abstract: Dusting roots of Abies concolor, Abies magnifica var. shastensis, Pseudotsuga menziesii and Pinus ponderosa with Pisolithus tinctorius (Pt) spores when planted out produced no Pt mycorrhizae at the end of the first growing season. In the 3rd yr occasional Pt mycorrhizae had formed on A. concolor. Inoculations reduced seedling survival in some cases. High rates of spore application may have desiccated roots of the true firs and spore amounts applied need careful attention. Soil scarification and ripping significantly promoted growth of A. concolor seedlings compared with scarification alone.

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8.
Amaranthus, M.P. and D.A. Perry. 1987. Effect of soil transfer on ectomycorrhiza formation and the survival and growth of conifer seedlings on old, nonreforested clear-cuts. Canadian-Journal-of-Forest-Research 17(8): 944-950.

Keywords:      planting operations

                        tree/stand health

                        growth

                        mycorrhizal response

Abstract: Small amounts (150 ml) of soil from established conifer plantations and mature forest were transferred to planting holes on 3 sites in the Klamath Mts., S. Oregon and N. California. The sites had been clear felled and burned 8-27 yr earlier and unsuccessfully reforested. At Cedar Camp, a high alt. (1720 m) southerly slope with sandy soil, transfer of soil from a Douglas fir plantation increased first-yr survival of Douglas fir seedlings by 50%, mycorrhizal formation and b.a. growth. Soil from mature forest did not enhance survival and growth. Soil transfer was less effective on 2 sites at lower alt. with clayey soils. Douglas fir seedlings at Crazy Peak showed similar, but less well defined, patterns to those at Cedar Camp. All Pinus lambertiana seedlings at Wood Creek survived well and were generally unaffected by soil transfer. Results suggest that adequate mycorrhizal formation is critical to seedling growth and survival on cold, droughty sites. Transfer of soil from a suitable source may offset the decline in native mycorrhizal fungi if reforestation is delayed.

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9.
Amaranthus, M.P. and D.A. Perry. 1989a. Interaction effects of vegetation type and Pacific madrone soil inocula on survival, growth and mycorrhiza formation of Douglas-fir. Canadian-Journal-of-Forest-Research 19(5): 550-556.

Keywords:      planting operations

                        growth

                        tree/stand health

                        mycorrhizal response

Abstract: One-yr-old non-mycorrhizal Douglas fir (Pseudotsuga menziesii) seedlings were planted in 1985 in cleared blocks within 3 adjacent vegetation types in SW Oregon, viz., whiteleaf manzanita (Arctostaphylos viscida), annual grass meadow, and an open stand of Oregon white oak (Quercus garryana). Within subplots in each block, either pasteurized or unpasteurized soil from a nearby Pacific madrone (Arbutus menziesii) stand was transferred to the planting holes of the seedlings; control seedlings received no madrone soil. Second-year survival averaged 92, 43 and 12% for seedlings planted on the manzanita, meadow and oak sites, respectively. Growth differences generally paralleled survival differences. Added madrone soil, whether pasteurized or unpasteurized, did not influence survival. Unpasteurized madrone soil substantially increased the growth of seedlings on the manzanita site, but not in the meadow or oak stand. Pasteurized madrone soil did not affect growth in any of the vegetation types. Unpasteurized madrone soil nearly tripled the number of mycorrhizal root tips forming on seedlings and resulted in formation of a new mycorrhiza type on the manzanita site, although it had little or no effect on the meadow or oak sites. These results suggest that manzanita and madrone impose a biological pattern on soils that stimulates Douglas fir growth and survival, and support results of other studies indicating that root symbionts and rhizosphere organisms mediate interactions among plant species.

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10.
Amaranthus, M.P. and D.A. Perry. 1989b. Rapid root tip and mycorrhiza formation and increased survival of Douglas-fir seedlings after soil transfer. New-Forests 3(3): 259-264.

Keywords:      planting operations

                        mycorrhizal response

                        root development

                        tree/stand health

Abstract: In order to re-inoculate soil with mycorrhizal fungi, small amounts (about 150 ml) of soil from an established Douglas fir (Pseudotsuga menziesii) plantation were added to planting holes when Douglas fir seedlings were planted on an old, unrevegetated clearcut in the Klamath Mountains of Oregon. Seedlings were lifted throughout the growing season to determine the influence of soil transfer on the rate of root tip initiation and mycorrhiza formation. Six weeks after planting, seedlings receiving plantation soil had formed 62% more root tips than controls; however, no statistically significant differences were apparent 15 weeks after planting. By that time, a small percentage of root tips were visibly mycorrhizal; seedlings receiving transferred soil had the most colonization (13.6 vs. 3.5 per seedling, p <less or =>0.05). Of seedlings receiving transfer soil, 36.6% survived the first growing season, compared to 11.3% of control seedlings. At this high altitude, soils often remain frozen well into spring, leaving only a brief period between the time when soils become warm enough for root growth and the onset of summer drought. Under these conditions, the rapid root growth and mycorrhiza formation stimulated by plantation soil increases the ability of seedlings to survive the first growing season.

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11.
Anekonda, T.S., M.C. Lomas, W.T. Adams, K.L. Kavanagh and S.N. Aitken. 2002. Genetic variation in drought hardiness of coastal Douglas-fir seedlings from British Columbia. Canadian-Journal-of-Forest-Research 32(10): 1701-1716.

Keywords:      genetic tree improvement

                        tree/stand protection

                        growth

                        tree/stand health

                        genetic relationships

                        tree physiology

Abstract: Genetic variation in drought hardiness traits and their genetic correlations with growth potential and recovery traits were investigated in 39 full-sib families of coastal Douglas-fir (Pseudotsuga menziesii var. menziesii) from southwestern British Columbia, Canada. Seedlings of these families were grown in raised nursery beds and subjected to three moisture regimes each in the second (well-watered or control, mild, and moderate drought) and third (control, severe drought, and recovery from second-year moderate drought) seasons. Traits assessed included drought hardiness (foliage damage, cavitation of xylem tracheids, xylem hydraulic conductivity, and height and diameter growth increment) in the drought treatments, growth potential (total height and diameter) in the control treatment, and height and diameter growth increments in the recovery treatment. Xylem cavitation in the growth ring produced in a particular year was nearly three times greater under the moderate drought and four times greater under the severe drought than in the control treatment. Xylem hydraulic conductivity of seedlings in the severe drought treatment was 40% lower than conductivity of seedlings under the control treatment. Mean foliage damage in seedlings subjected to severe drought (third season) was much greater (33%) than in seedlings subjected to mild or moderate drought (second season). Families differed significantly in most drought hardiness traits, with individual tree heritabilities averaging 0.19. Thus, much potential exists for identifying drought-hardy families at the seedling stage and using this information for deployment or breeding purposes. In addition, most hardiness traits were strongly intercorrelated (genetic correlations often exceeded |0.80|) indicating that these traits are controlled largely by the same set of genes and that selection for hardiness based on one trait will increase hardiness as reflected in the other traits as well. Genetic correlations were only moderate (0.49) between hardiness traits measured in different years, perhaps due to the large difference in severity of the drought applied in the two seasons. Although injury to seedlings, as reflected in foliage damage and xylem cavitation, was relatively low under the moderate drought of the second season, it did result in reduced growth increment the following (recovery) year. Growth potential under favourable moisture regimes was nearly uncorrelated with drought hardiness, suggesting that drought hardiness could be improved in this southwestern British Columbia breeding population without negatively impacting growth potential in favourable moisture conditions.

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12.
Antonelli, A.L. and R.L. Campbell. 1991. Cooley spruce gall aphid. College of Agriculture and Home Economics, Washington State University. Extension Bulletin EB0966: 2 p.

Keywords:      tree/stand protection

                        tree/stand health

Abstract: Notes are provided on the biology, injuriousness and chemical control (carbaryl and endosulfan are suggested) of Adelges cooleyi [Gilletteella cooleyi] on certain coniferous trees [including Picea sitchensis, P. engelmannii, P. pungens and Pseudotsuga menziesii] in Washington State.

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13.
Arnott, J.T. and D. Beddows. 1982. Influence of Styroblock container size on field performance of Douglas-fir, western hemlock, and Sitka spruce. Tree Planters' Notes 33(3): 31-34.

Keywords:      nursery operations

                        growth

                        tree/stand health

Abstract: Pseudotsuga menziesii, Tsuga heterophylla and Picea sitchensis seeds were sown in April 1971 in BC/CFS Styroblocks sizes 2 and 8 with volumes of 40 and 125 cmsuperscript 3 respectively. The seedlings in the larger containers were kept in a heated greenhouse for 2-3 months to stimulate growth to fill the containers, before joining those in the smaller containers in an outdoor shadehouse nursery. Seedlings were planted out in British Columbia in March 1972. A second trial was started in April 1972 and seedlings planted out in April 1973. Survival and ht. growth were recorded for 5 growing seasons. The larger containers produced larger seedlings at planting. There were n.s.d. in survival of seedlings grown in the different sized containers for all 3 species. The seedlings grown in the larger containers were significantly taller after the first growing season in the field, a difference which persisted for the 5 seasons. Growing seedlings in the larger containers was more expensive and the seedlings took longer to plant than those grown in the smaller containers.

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14.
Arnott, J.T. and F.T. Pendl. 1994. Field performance of several tree species and stock types planted in montane forests of coastal British Columbia. Canadian-Forest-Service, Pacific and Yukon Region Information Report BC-X-347. viii + 45 p.

Keywords:      nursery operations

                        planting operations

                        growth

                        tree/stand health

                        wood quality

Abstract: Planting trials were established at sites within the Mountain Hemlock and montane Coastal Western Hemlock biogeoclimatic zones. Six test areas were chosen within each zone. Amabilis fir (Abies amabilis), noble fir (A. procera), yellow cedar (Chamaecyparis nootkatensis) and mountain hemlock (Tsuga mertensiana) were the species selected for planting in the Mountain Hemlock zone. In addition to Abies amabilis and A. procera, western white pine (Pinus monticola), western redcedar (Thuja plicata), Douglas fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) were planted in the Coastal Western Hemlock zone. Plug (PSB 211), plug transplant and bareroot stock types were used for the eight species across both zones. Seedlings were planted during the autumn (September/October) and spring (May) in each of two successive years: 1978-79 and 1979-80. Survival, growth and tree form 13 years after planting were used as indicators of the reliability (a combination of tree survival and form) and productivity of the planting treatment combinations. Noble fir and amabilis fir were the most reliable species in the Mountain Hemlock zone; i.e. these species have average survival rates higher than 80% and few form defects. Yellow cedar crowns were badly broken by snow, which reduced the reliability of this species in the early years of plantation establishment. The growth, survival and form of mountain hemlock ranked between that of the true firs and yellow cedar. Noble fir was by far the most productive species in the Mountain Hemlock zone. Within the Coastal Western Hemlock zone no single species demonstrated a superior combination of productivity or reliability. Douglas fir, western hemlock and western redcedar were good species in the lower elevations of the zone, whereas noble fir and amabilis fir were better species at the upper elevational limits of the zone. Western white pines should be avoided until rust-resistant seed sources are available. Little variation was found among the three planting stock options and even less between the two planting seasons. Plug transplant stock was more reliable than bareroot or plug stock; productivity ranked from greatest to least in the following order within both zones: plug transplant, bareroot and plug stock. This ranking among stock types may well change as different stock types are developed. However, the relative size and design differences among stock types, no matter when they become available, will always have an effect on the ultimate reliability and productivity of planted trees. Autumn planting gave significantly lower survival in the Coastal Western Hemlock zone only.

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15.
Axelrood, P.E., W.K. Chapman, K.A. Seifert, D.B. Trotter and G. Shrimpton. 1998. Cylindrocarpon and Fusarium root colonization of Douglas-fir seedlings from British Columbia reforestation sites. Canadian Journal of Forest Research 28:1198-1206.

Keywords:      planting operations

                        tree/stand protection

                        tree/stand health

Abstract: Poor performance of Douglas fir (Pseudotsuga menziesii) plantations established in 1987 has occurred in southwestern British Columbia. Affected sites were planted with 1-yr-old container stock that exhibited some root dieback in the nursery. A study was initiated in 1991 to assess Cylindrocarpon and Fusarium root infection in planted and naturally regenerating (natural) Douglas fir seedlings from 7 affected plantations. Percentages of seedlings harbouring Cylindrocarpon spp. and percentage root colonization were significantly greater for planted seedlings than natural seedlings. A significant linear trend in Cylindrocarpon root colonization was observed for planted seedlings with colonization levels being highest for roots closest to the remnants of the root plug and decreasing at distances greater than 10 cm from that region. This trend in Cylindrocarpon colonization was not observed for natural seedlings. Cylindrocarpon destructans var. destructans [Nectria radicola var., radicola] and C. cylindroides var. cylindroides were the only species isolated from planted and natural conifer seedlings. For most sites, percentage of seedlings harbouring Fusarium spp. and percentage Fusarium root colonization were less than for Cylindrocarpon. Recovery of Fusarium spp. from seedlings and root colonization levels were not significantly different for planted and natural seedlings from all sites.

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16.
Axelrood, P.E., M. Neumann, D. Trotter, R. Radley, G. Shrimpton and J. Dennis. 1995. Seedborne Fusarium on Douglas-fir: pathogenicity and seed stratification method to decrease Fusarium contamination. New-Forests 9(1): 35-51.

Keywords:      nursery operations

                        tree/stand protection

                        tree/stand health

                        growth

Abstract: Twelve Douglas-fir (Pseudotsuga menziesii) seedlots from coastal British Columbia were assayed for seedborne Fusarium; all of the seedlots were contaminated. The percentage of non-stratified seeds from individual seedlots harbouring Fusarium ranged from 0.3 to 95.4. Sixty-seven percent of the seedlots had Fusarium on less than 2% of the seeds. Post-stratification seedborne Fusarium levels were significantly less for running water imbibition compared with standing water imbibition. However, seedling growth at a container nursery was not consistently different for stratified seed imbibed initially in standing or running water. Fusarium disease symptoms were not observed in the nursery environment. The species of Fusarium isolated from seed were F. acuminatum [Gibberella acuminata], F. avenaceum [G. avenacea], F. lateritium [G. baccata], F. moniliforme [G. fujikuroi], F. oxysporum, F. poae and F. sambucinum [G. pulicaris]. Twelve Fusarium isolates, comprising 6 species, were assessed for pathogenicity. Disease symptoms were observed after 4 weeks incubation and Fusarium isolates ranged in virulence from low to high. Fusarium oxysporum isolates were the most pathogenic.

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17.
Axelrood, P.E. and R. Radley. 1991. Biological control of Fusarium on Douglas-fir seedlings. Bulletin-SROP 14(8): 85-87.

Keywords:      nursery operations

                        tree/stand protection

                        tree/stand health

Abstract: A bacterial culture collection was established from the rhizosphere and rhizoplane of Douglas fir (Pseudotsuga menziesii) seedlings collected from nursery and forest locations in British Columbia, Canada. Of the 2000 strains screened, 350 inhibited growth of at least 1 conifer seedling root pathogen (Fusarium, Cylindrocarpon or Pythium) in in vitro antibiosis assays. A total of 96 strains were screened for Fusarium disease control in biological control assays. One strain that inhibited all 3 pathogens in vitro was able to significantly reduce the incidence of disease caused by Fusarium on P. menziesii seedlings. Another strain that tested negative in in vitro antibiosis assays also reduced the disease incidence by a similar amount. This paper was presented at the Second international workshop on plant growth-promoting rhizobacteria - progress and prospects, held in Interlaken, Switzerland, Oct. 14-19, 1990.


18.
Barclay, H.J. and H. Brix. 1984. Effects of urea and ammonium nitrate fertilizer on growth of a young thinned and unthinned Douglas-fir stand. Canadian-Journal-of-Forest-Research 14(6): 952-955.

Keywords:      fertilization

                        thinning

                        growth

                        tree physiology

                        tree/stand health

Abstract: The effects were studied of 2 sources of nitrogen fertilizer applied at rates of 224 and 448 kg/ha N on growth of thinned and unthinned plots established in 1970 in a 24-yr-old stand on southern Vancouver Is., British Columbia. Ammonium nitrate yielded higher growth of diam. and vol. than urea over a 9-yr period, particularly with thinning. Ht. growth was not affected by nitrogen source. The efficiency of nitrogen fertilizing in terms of stem vol. response per kilogram of nitrogen applied was greatest with ammonium nitrate in thinned plots. Tree mortality increased substantially with fertilizing for both sources, and decreased markedly with thinning.

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19.
Barclay, H.J. and H. Brix. 1985a. Effects of high levels of fertilization with urea on growth of thinned and unthinned Douglas-fir stands. Canadian-Journal-of-Forest-Research 15(4): 730-733.

Keywords:      fertilization

                        thinning

                        growth

                        tree physiology

                        tree/stand health

Abstract: Diameter height and volume growth were documented for 9 yr after thinning and fertilizing in a 24-yr-old stand on a poor site on southern Vancouver Is. The treatments involved 3 thinning treatments (0, 1/3, and 2/3 b.a. removed) and 6 fertilizer treatments (0-1344 kg/ha N) with urea. Increments for both diameter and gross volume increased with the rate of fertilizer application and responses were still apparent 9 yr after treatment. For unthinned plots, the 9-yr volume growth responses were 30, 50, and 80% with fertilizer rates of 224, 448, and 896 kg/ha N, respectively. The efficiency of fertilizer use, measured as stem volume response per unit of nitrogen applied, decreased with rate of fertilizer application, but this result may change over a longer response period. There was a positive interaction between fertilizing and thinning such that high amounts of both mutually enhanced growth. Mortality increased with fertilizing, but only noticeably in unthinned plots.

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20.
Barclay, H.J. and C.R. Layton. 1990. Growth and mortality in managed Douglas fir: relation to a competition index. Forest-Ecology-and-Management 36(2-4): 187-204.

Keywords:      fertilization

                        thinning

                        growth

                        tree/stand health

Abstract: Twelve-year increments of diameter at breast height (DBH), height, and volume in thinned and fertilized 45-year-old Douglas fir (Pseudotsuga menziesii) stands on Vancouver Island, Canada, were related (by regression) to degree of thinning, amount of fertilizer (3x3 factorial), initial DBH, and a competitive stress index (CSI). The ability of the CSI to predict growth after treatment was examined. Causes of tree death, and CSI data, are presented, and the relationship between them discussed. The CSI was found to be only moderately good at predicting Douglas fir growth and mortality: initial DBH provided a better predictor. Most mortality in unthinned plots resulted from suppression, and correlated reasonably well with CSI; mortality in thinned plots was not correlated with CSI, and resulted principally from snow damage. Tree height variability generally became less over the 12 years following treatment, which is more consistent with two-sided than one-sided competition predictions, a result which is contrasted to that of many other species.

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21.
Belz, D. and T.E. Nishimura. 1989. Effects of imazapyr, 2,4-D and metsulfuron methyl on conifer tolerance. Proceedings-of-the-Western-Society-of-Weed-Science (Vol. 42): 98-104.

Keywords:      site preparation

                        chemical preparation

                        release treatments

                        chemical release

                        tree/stand health

Abstract: Imazapyr at 0.25-1.0 lb/acre alone or 0.5 lb/acre in combination with 2,4-D 2 lb/acre or metsulfuron 0.3 lb/acre was evaluated for effect on growth and injury to Pseudotsuga menziesii, Pinus ponderosa, Tsuga heterophylla and Abies amabilis seedlings in the Pacific Northwest region. Applications were made at 4 times: 3 month pre-planting in Dec., as buds began to swell in Mar., during the spring flush of growth in May, and after bud set in Aug. The effect of different application rates was of less significance than their timings. Application during active growth gave unacceptable injury levels; pre-planting caused least injury, but autumn treatment was acceptable for tolerant species. Species tolerance was in the order Pinus ponderosa > Pseudotsuga menziesii > T. heterophylla > A. amabilis.

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22.
Bettinger, P., K.A. Bettinger and K. Boston. 1998. Correlation among spatial and non-spatial variables describing a cut-to-length thinning site in the Pacific Northwest, USA. Forest-Ecology-and-Management 104(1/3): 139-149.

Keywords:      thinning

                        commercial thinning

                        tree/stand health

Abstract: Variables describing the pre- and post-logging conditions of a thinning site in 47-yr-old naturally regenerated stand of second-growth Douglas fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) in western Oregon, were examined for correlation, and subsequently used to develop models to estimate residual stand damage levels. A cut-to-length harvesting system was utilized to perform the thinning operation, which used a single-grip harvester and a forwarder, and marked logging trails. Several of the variables were measured in an intensive field survey; other variables were developed using geographic information system (GIS) processes. An analysis of correlations among the site variables showed several obvious, and a few interesting, results that describe the operation. Most of the variables provided negative, or inconclusive, assistance in describing the variation in stand damage levels. Only one variable, the number of original trees/hectare, was significantly correlated with residual stand damage levels, and was represented in the models that were developed to estimate residual stand damage levels. The resulting models are of limited practical value, however, since they explain little of the variability in damage levels. Most of the variation in residual stand damage levels may well be explained by random chance, operator error, other unmeasured operational variables associated with this harvesting system, or interactions among variables. The main conclusion from the study is that although both spatial and non-spatial data were utilized in describing the logging operation and in developing models to estimate stand damage levels, the importance of using spatial data was inconclusive.

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23.
Bettinger, P. and L.D. Kellogg. 1993. Residual stand damage from cut-to-length thinning of second-growth timber in the Cascade Range of western Oregon. Forest-Products-Journal 43(11/12): 59-64.

Keywords:      thinning

                        commercial thinning

                        tree/stand health

Abstract: Residual stand damage was measured on 25% of an area that had been thinned with a cut-to-length logging system. Total damage (scar area) per acre was less than in any similar study in the Pacific Northwest, although 39.8% of the residual trees sustained some damage. Only 0.8% of the trees, however, sustained significant damage. Western hemlock (Tsuga heterophylla) was more susceptible to damage than Douglas fir (Pseudotsuga menziesii). Most of the damage occurred within 15 feet of a trail centreline and originated within 3 feet of the groundline. Early summer logging may have resulted in more damaged trees than might occur during other seasons. Future volume loss due to decay is likely to be minimal because a low percentage of scars were considered vulnerable to wood-decaying fungi.

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24.
Binkley, D. 1984. Importance of size-density relationships in mixed stands of Douglas-fir and red alder. Forest-Ecology-and-Management 9(2): 81-85.

Keywords:      thinning

                        growth

                        tree/stand health

Abstract: Pairs of Douglas-fir, and Douglas-fir and red alder (Alnus rubra) stands were examined at four locations (in Oregon, Washington and British Columbia) for patterns in average tree size as a function of stand density. On fertile sites, the mixed stands experienced higher mortality than the pure conifer stands. On infertile sites, the pure conifer stands were well below the maximum tree size and density relationship compared to fertile sites or mixed stands, suggesting under-utilized site resources were available for nitrogen-fixing alder.

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25.
Birchler, T.M., R. Rose and D.L. Haase. 2001. Fall fertilization with N and K: effects on Douglas-fir seedling quality and performance. Western-Journal-of-Applied-Forestry 16(2): 71-79.

Keywords:      nursery operations

                        nursery fertilization

                        growth

                        tree physiology

                        tree morphology

                        tree/stand health

Abstract: Coastal Douglas fir (Pseudotsuga menziesii) 1+1 seedlings from coastal Oregon, USA, were applied with two fertilizers (NH4NO3+K2SO4 and (NH4)2SO4+KCl) at four rates (0, 80, 160, 320 kg N and K/ha) split over three application dates (September 19, October 13, November 1, 1996). Fertilizer type did not affect total Kjeldahl nitrogen (TKN) levels on any of the sampling dates. By January 10, TKN concentrations had increased 16, 30 and 34%, and chloride concentrations had increased 57, 77 and 112% relative to the seedlings without fertilizer, for 80, 160 and 320 kg N+K/ha treatments, respectively. Nitrate levels increased briefly after the first application of NH4NO3+K2SO4. Potassium levels remained relatively unchanged. Levels of most other nutrients, as well as foliar dry weight, increased between September 16 and January 10, but these increases were generally unrelated to the fertilizer treatments. Root growth potential and cold hardiness did not differ among treatments. Seedlings that received 160 or 320 kg N/ha broke bud an average of 3 days earlier than the seedlings without fertilizer. Chlorophyll fluorescence (Fv/Fm) of seedlings with fertilizer was consistently higher than that of seedlings without fertilizer on November 13 and December 30. These treatment differences were not reflected in seedling outplanting performance after one growing season.

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26.
Blake, J.I., R. Linderman and D. Lavender. 1988b. Seedling vigor of Douglas fir and western hemlock in relation to ethylene exposure levels and ethane production during cold storage. In Proceedings: 10th North American Forest Biology Workshop, 'Physiology and genetics of reforestation', University of British Columbia, Vancouver, British Columbia, July 10-22, 1988. Eds. J. Worrall, J. Loo-Dinkins and D.P. Lester. pp. 235-242.

Keywords:      nursery operations

                        tree morphology

                        tree phenology

                        tree/stand health

Abstract: The effects were examined of ethylene treatment on Pseudotsuga menziesii and Tsuga heterophylla 2-yr-old bare root seedlings lifted in October or December (in a nursery in Washington) and stored for 2 and 7 days. Seedlings exposed to 0.5 p.p.m. ethylene at +1 degrees C for 7 days exhibited reduced bud and root activity. These effects were apparently reversed at 5.0 p.p.m. In a second experiment, reducing ethylene concentrations with KMnO4 during storage at +1 or +10 degrees C for 30 days did not affect foliage colour, but root and bud activity were generally enhanced. Ethylene concentrations in control bags ranged from 0.80 to 2.24 p.p.m. in October-lifted seedling bags and from 0.10 to 1.3 p.p.m. in December-lifted samples. The quantity of ethane in stored bags of P. menziesii seedlings was closely related to an increase in foliage discoloration. Little or no ethane was measured in T. heterophylla storage bags.

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27.
Blake, J.I. and R.G. Linderman. 1992. A note on root development, bud activity, and survival of Douglas-fir, and survival of western hemlock and noble-fir seedlings, following exposure to ethylene during cold storage. Canadian-Journal-of-Forest-Research 22(8): 1195-1200.

Keywords:      nursery operations

                        tree morphology

                        tree phenology

                        tree/stand health

Abstract: Three cold storage experiments were conducted with bare-root (2+0) Douglas fir (Pseudotsuga menziesii) seedlings from coastal Oregon and eastern Washington Cascade sources. The objectives were to determine the effects of ethylene at ambient and below-ambient (absorbed by KMnO4 pellets) concentrations, and at 0.5 and 5 p.p.m. ethylene, during short-term storage on subsequent root development and bud activity, and to relate these results to survival in the field at sites in Washington and Oregon, after prolonged cold storage. Root numbers and lengths were measured 28 days after a 7-day storage period after lifting seedlings on 27 September and 1 December. In the coastal source, root numbers and lengths in the 5 p.p.m. ethylene treatment were, respectively, 46 and 49% greater in September, and 22 and 13% greater in December, than the controls. No comparable treatment effects were found for the Cascade source. Neither the KMnO4 nor the 0.5 p.p.m. ethylene treatments affected root development in either seed source. For terminal buds in the controls, the number of days to 50% bud break was increased 2-8 days by a 30-day cold storage period compared with a 7-day period. For the coastal source, no increase in the time to 50% bud break was observed in the 5 p.p.m. ethylene treatment. Seedling survival was evaluated in the field for the same treatments following 4 months cold storage for the Douglas fir sources, coastal western hemlock (Tsuga heterophylla), and noble fir (Abies procera). Survival for the 5 p.p.m. ethylene treatment compared with the control was increased by 55% in the coastal Douglas fir source and by 13% in western hemlock. These results suggest that stimulated root development and bud activity may be partially responsible for the observed survival increase following cold storage at elevated ethylene levels.

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28.
Bledsoe, C.S. and R.J. Zasoski. 1983. Effects of ammonium and nitrate on growth and nitrogen uptake by mycorrhizal Douglas-fir seedlings. In Tree root systems and their mycorrhizas. Ed. D. Atkinson. pp. 445-454.

Keywords:      nursery operations

                        nursery fertilization

                        growth

                        tree physiology

                        tree morphology

                        tree/stand health

Abstract: In a greenhouse pot study, 1-yr-old mycorrhizal (inoculated with Hebeloma crustuliniforme) and non-mycorrhizal Douglas fir seedlings were grown in sandy forest soil amended with 10% of clay minerals (bentonite and/or kaolinite) and ammonium or nitrate fertilizer. Ht. growth, root and shoot DM and accumulation of nitrogen and P were greater in mycorrhizal than non-mycorrhizal seedlings, especially in the nitrate treatment. Ammonium interacted with kaolinite to reduce survival which again was poorer in the absence of mycorrhiza.

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29.
Bloomberg, W.J. 1988. Modeling control strategies for laminated root rot in managed Douglas-fir stands: model development. Phytopathology 78(4): 403-409.

Keywords:      planting operations

                        site preparation

                        mechanical preparation

                        tree/stand protection

                        tree/stand health

Abstract: A model of laminated root rot caused by Phellinus [Inonotus] weirii was developed to assess potential control strategies in managed Pseudotsuga menziesii stands. The model mimicked key processes in disease initiation and development quantified as functions of time and space. These processes were horizontal and vertical tree root distribution, root contact with inoculum and among root systems, spread of mycelium through root systems, root decay, reduction of diam. growth in infected trees, tree mortality and persistence of inoculum in roots of stumps and killed trees. The processes were expressed as mathematical functions which were integrated in a computer program to calculate spread of the disease and stand-growth loss and mortality. Data for quantification of functions were obtained by experiments and from the literature. Simulated control practices included infected stump removal, sanitation fellings and mixed planting of Douglas fir and resistant species. Accuracy of the model was tested by comparing calculated disease spread and mortality with the following data: (1) spread and damage in two 60-yr-old, 1-ha stands in Oregon, (2) results from a statistically based model for spread and damage that had performed satisfactorily, and (3) observed spread and damage behaviour in stands of different ages and growth rates. Results from the model compared favourably with all of the above situations.

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30.
Bloomberg, W.J. and G. Reynolds. 1988. Equipment trials for uprooting root-rot-infected stumps. Western-Journal-of-Applied-Forestry 3(3): 80-82.

Keywords:      site preparation

                        mechanical preparation

                        tree/stand protection

                        tree/stand health

Abstract: Residual roots from Douglas fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) were measured following stump-root extraction one yr after harvesting a 55-yr-old, 314 stems/ha, 47% Douglas fir, 17% maple (Acer macrophyllum), 16% red cedar (Thuja plicata), 6% western hemlock stand with 20% infection by Phellinus weirii in the Cowichan valley, Vancouver Island, Canada. Extraction was by a Caterpillar D8H with brush-clearing blade, a 180-hp backhoe or a 115-hp backhoe. All 3 machines recovered more than 90% of root vol. The small backhoe left significantly greater numbers and lengths of root residues per msuperscript 3 soil, though the vol. of residues was greatest for the Caterpillar. An earlier study suggested that a root density of 32 roots/msuperscript 3 was needed to produce one root contact; as the least efficient treatment by the Caterpillar left 23.2 roots/msuperscript 3 in the ground, it is suggested that this would provide insufficient contacts with a new tree crop to transmit infection.

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31.
Brand, D.G. 1986a. A competition index for predicting the vigour of planted Douglas-fir in southwestern British Columbia. Canadian-Journal-of-Forest-Research 16(1): 23-29.

Keywords:      planting operations

                        tree/stand health

                        growth

Abstract: As a method of quantifying brush competition, data from 124 planted Douglas firs, age 1-5 yr, were used to derive a competition index to predict changes in tree vigour measured as a relative production rate. The index, which includes measures of brush proximity, relative ht. and % ground cover, appears to act as a measure of light interception around the tree crown. Tree vigour was found to be largely a function of the age of the tree from planting and the competition index. Foliage-based measures of growth vigour were related more strongly to the index than mea