1. Adams, W.T., J. Zuo, J.Y. Shimizu, J.C. Tappeiner and J.H. Zuo. 1998. Impact of alternative regeneration methods on genetic diversity in coastal Douglas-fir. Forest Science 44:390-396.

Keywords:      planting operations

                        genetic relationships

Abstract: Genetic markers (17 allozyme loci) were used to study the genetic implications of natural and artificial regeneration following 3 regeneration methods (group selection, shelterwood, and clearcut) in coastal Douglas fir (Pseudotsuga menziesii var. menziesii) forests in the Willamette Valley, Oregon, USA. In general, harvesting followed by either natural or artificial regeneration resulted in offspring populations which were very similar to those in the previous generation. The removal of the smallest trees to form shelterwoods resulted in the removal of rare, presumably deleterious, alleles. Fewer alleles per locus were observed among residual trees (2.76) and natural regeneration (2.75) than were found in uncut (control) stands (2.86). The shelterwood regime appeared quite compatible with gene conservation, and it was considered that it was best to leave parent trees of a range of sizes to act as gene conservation reserves, in order to maximize the number of alleles (regardless of current adaptive value) in naturally regenerated offspring. Seedling stocks used for artificial regeneration in clearcut, shelterwood, and group selection stands (7 total) had significantly greater levels of genetic diversity, on average, than natural regenerated seedlings.

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2.
Alvarez, I.F. and J.M. Trappe. 1983a. Dusting roots of Abies concolor and other conifers with Pisolithus tinctorius spores at outplanting time proves ineffective. Canadian-Journal-of-Forest-Research 13(5): 1021-1023.

Keywords:      planting operations

                        site preparation

                        mechanical preparation

                        growth

                        tree/stand health

                        mycorrhizal response

Abstract: Dusting roots of Abies concolor, Abies magnifica var. shastensis, Pseudotsuga menziesii and Pinus ponderosa with Pisolithus tinctorius (Pt) spores when planted out produced no Pt mycorrhizae at the end of the first growing season. In the 3rd yr occasional Pt mycorrhizae had formed on A. concolor. Inoculations reduced seedling survival in some cases. High rates of spore application may have desiccated roots of the true firs and spore amounts applied need careful attention. Soil scarification and ripping significantly promoted growth of A. concolor seedlings compared with scarification alone.

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3.
Amaranthus, M.P., C.Y. Li and D.A. Perry. 1990. Influence of vegetation type and madrone soil inoculum on associative nitrogen fixation in Douglas-fir rhizospheres. Canadian-Journal-of-Forest-Research 20(3): 368-371.

Keywords:      planting operations

                        soil properties

Abstract: In studies in SW Oregon in 1985, Douglas fir (Pseudotsuga menziesii) seedlings grown at a site cleared of whiteleaf manzanita (Arctostaphylos viscida) and an adjacent, cleared, annual grass meadow were either inoculated with 100-120 ml per seedling of pasteurized or unpasteurized soil from a nearby Pacific madrone (Arbutus menziesii) stand, or left uninoculated. After one growing season, Douglas fir seedling whole-plant soil systems were assayed for nitrogenase activity by the acetylene reduction method. The rate of acetylene reduction in rhizospheres of uninoculated seedlings from the manzanita site (1.40 +or- 0.44 nmol/h) was significantly greater than that of uninoculated seedlings from the meadow site (0.67 +or- 0.15 nmol/h). Unpasteurized madrone soil increased the rate of acetylene reduction by >500% for inoculated seedlings grown on the manzanita site, but decreased it by 80% for those grown on the meadow site. The influence of madrone soil was apparently biotic: pasteurized madrone soil did not have a significant effect. No acetylene was reduced in soil without seedlings. Azospirillum sp., a microaerophilic nitrogen (N2) fixing bacterium, was isolated from within the mycorrhizas of inoculated seedlings harvested from the manzanita site. These results suggest that early successional ectomycorrhizal shrubs and broadleaved trees may be important in maintaining mycorrhizal fungi and associated N2 fixers after severe disturbance.

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4.
Amaranthus, M.P. and D.A. Perry. 1987. Effect of soil transfer on ectomycorrhiza formation and the survival and growth of conifer seedlings on old, nonreforested clear-cuts. Canadian-Journal-of-Forest-Research 17(8): 944-950.

Keywords:      planting operations

                        tree/stand health

                        growth

                        mycorrhizal response

Abstract: Small amounts (150 ml) of soil from established conifer plantations and mature forest were transferred to planting holes on 3 sites in the Klamath Mts., S. Oregon and N. California. The sites had been clear felled and burned 8-27 yr earlier and unsuccessfully reforested. At Cedar Camp, a high alt. (1720 m) southerly slope with sandy soil, transfer of soil from a Douglas fir plantation increased first-yr survival of Douglas fir seedlings by 50%, mycorrhizal formation and b.a. growth. Soil from mature forest did not enhance survival and growth. Soil transfer was less effective on 2 sites at lower alt. with clayey soils. Douglas fir seedlings at Crazy Peak showed similar, but less well defined, patterns to those at Cedar Camp. All Pinus lambertiana seedlings at Wood Creek survived well and were generally unaffected by soil transfer. Results suggest that adequate mycorrhizal formation is critical to seedling growth and survival on cold, droughty sites. Transfer of soil from a suitable source may offset the decline in native mycorrhizal fungi if reforestation is delayed.

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5.
Amaranthus, M.P. and D.A. Perry. 1989a. Interaction effects of vegetation type and Pacific madrone soil inocula on survival, growth and mycorrhiza formation of Douglas-fir. Canadian-Journal-of-Forest-Research 19(5): 550-556.

Keywords:      planting operations

                        growth

                        tree/stand health

                        mycorrhizal response

Abstract: One-yr-old non-mycorrhizal Douglas fir (Pseudotsuga menziesii) seedlings were planted in 1985 in cleared blocks within 3 adjacent vegetation types in SW Oregon, viz., whiteleaf manzanita (Arctostaphylos viscida), annual grass meadow, and an open stand of Oregon white oak (Quercus garryana). Within subplots in each block, either pasteurized or unpasteurized soil from a nearby Pacific madrone (Arbutus menziesii) stand was transferred to the planting holes of the seedlings; control seedlings received no madrone soil. Second-year survival averaged 92, 43 and 12% for seedlings planted on the manzanita, meadow and oak sites, respectively. Growth differences generally paralleled survival differences. Added madrone soil, whether pasteurized or unpasteurized, did not influence survival. Unpasteurized madrone soil substantially increased the growth of seedlings on the manzanita site, but not in the meadow or oak stand. Pasteurized madrone soil did not affect growth in any of the vegetation types. Unpasteurized madrone soil nearly tripled the number of mycorrhizal root tips forming on seedlings and resulted in formation of a new mycorrhiza type on the manzanita site, although it had little or no effect on the meadow or oak sites. These results suggest that manzanita and madrone impose a biological pattern on soils that stimulates Douglas fir growth and survival, and support results of other studies indicating that root symbionts and rhizosphere organisms mediate interactions among plant species.

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6.
Amaranthus, M.P. and D.A. Perry. 1989b. Rapid root tip and mycorrhiza formation and increased survival of Douglas-fir seedlings after soil transfer. New-Forests 3(3): 259-264.

Keywords:      planting operations

                        mycorrhizal response

                        root development

                        tree/stand health

Abstract: In order to re-inoculate soil with mycorrhizal fungi, small amounts (about 150 ml) of soil from an established Douglas fir (Pseudotsuga menziesii) plantation were added to planting holes when Douglas fir seedlings were planted on an old, unrevegetated clearcut in the Klamath Mountains of Oregon. Seedlings were lifted throughout the growing season to determine the influence of soil transfer on the rate of root tip initiation and mycorrhiza formation. Six weeks after planting, seedlings receiving plantation soil had formed 62% more root tips than controls; however, no statistically significant differences were apparent 15 weeks after planting. By that time, a small percentage of root tips were visibly mycorrhizal; seedlings receiving transferred soil had the most colonization (13.6 vs. 3.5 per seedling, p <less or =>0.05). Of seedlings receiving transfer soil, 36.6% survived the first growing season, compared to 11.3% of control seedlings. At this high altitude, soils often remain frozen well into spring, leaving only a brief period between the time when soils become warm enough for root growth and the onset of summer drought. Under these conditions, the rapid root growth and mycorrhiza formation stimulated by plantation soil increases the ability of seedlings to survive the first growing season.

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7.
Arnott, J.T. and F.T. Pendl. 1994. Field performance of several tree species and stock types planted in montane forests of coastal British Columbia. Canadian-Forest-Service, Pacific and Yukon Region Information Report BC-X-347. viii + 45 p.

Keywords:      nursery operations

                        planting operations

                        growth

                        tree/stand health

                        wood quality

Abstract: Planting trials were established at sites within the Mountain Hemlock and montane Coastal Western Hemlock biogeoclimatic zones. Six test areas were chosen within each zone. Amabilis fir (Abies amabilis), noble fir (A. procera), yellow cedar (Chamaecyparis nootkatensis) and mountain hemlock (Tsuga mertensiana) were the species selected for planting in the Mountain Hemlock zone. In addition to Abies amabilis and A. procera, western white pine (Pinus monticola), western redcedar (Thuja plicata), Douglas fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) were planted in the Coastal Western Hemlock zone. Plug (PSB 211), plug transplant and bareroot stock types were used for the eight species across both zones. Seedlings were planted during the autumn (September/October) and spring (May) in each of two successive years: 1978-79 and 1979-80. Survival, growth and tree form 13 years after planting were used as indicators of the reliability (a combination of tree survival and form) and productivity of the planting treatment combinations. Noble fir and amabilis fir were the most reliable species in the Mountain Hemlock zone; i.e. these species have average survival rates higher than 80% and few form defects. Yellow cedar crowns were badly broken by snow, which reduced the reliability of this species in the early years of plantation establishment. The growth, survival and form of mountain hemlock ranked between that of the true firs and yellow cedar. Noble fir was by far the most productive species in the Mountain Hemlock zone. Within the Coastal Western Hemlock zone no single species demonstrated a superior combination of productivity or reliability. Douglas fir, western hemlock and western redcedar were good species in the lower elevations of the zone, whereas noble fir and amabilis fir were better species at the upper elevational limits of the zone. Western white pines should be avoided until rust-resistant seed sources are available. Little variation was found among the three planting stock options and even less between the two planting seasons. Plug transplant stock was more reliable than bareroot or plug stock; productivity ranked from greatest to least in the following order within both zones: plug transplant, bareroot and plug stock. This ranking among stock types may well change as different stock types are developed. However, the relative size and design differences among stock types, no matter when they become available, will always have an effect on the ultimate reliability and productivity of planted trees. Autumn planting gave significantly lower survival in the Coastal Western Hemlock zone only.

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8.
Axelrood, P.E., W.K. Chapman, K.A. Seifert, D.B. Trotter and G. Shrimpton. 1998. Cylindrocarpon and Fusarium root colonization of Douglas-fir seedlings from British Columbia reforestation sites. Canadian Journal of Forest Research 28:1198-1206.

Keywords:      planting operations

                        tree/stand protection

                        tree/stand health

Abstract: Poor performance of Douglas fir (Pseudotsuga menziesii) plantations established in 1987 has occurred in southwestern British Columbia. Affected sites were planted with 1-yr-old container stock that exhibited some root dieback in the nursery. A study was initiated in 1991 to assess Cylindrocarpon and Fusarium root infection in planted and naturally regenerating (natural) Douglas fir seedlings from 7 affected plantations. Percentages of seedlings harbouring Cylindrocarpon spp. and percentage root colonization were significantly greater for planted seedlings than natural seedlings. A significant linear trend in Cylindrocarpon root colonization was observed for planted seedlings with colonization levels being highest for roots closest to the remnants of the root plug and decreasing at distances greater than 10 cm from that region. This trend in Cylindrocarpon colonization was not observed for natural seedlings. Cylindrocarpon destructans var. destructans [Nectria radicola var., radicola] and C. cylindroides var. cylindroides were the only species isolated from planted and natural conifer seedlings. For most sites, percentage of seedlings harbouring Fusarium spp. and percentage Fusarium root colonization were less than for Cylindrocarpon. Recovery of Fusarium spp. from seedlings and root colonization levels were not significantly different for planted and natural seedlings from all sites.

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9.
Bloomberg, W.J. 1988. Modeling control strategies for laminated root rot in managed Douglas-fir stands: model development. Phytopathology 78(4): 403-409.

Keywords:      planting operations

                        site preparation

                        mechanical preparation

                        tree/stand protection

                        tree/stand health

Abstract: A model of laminated root rot caused by Phellinus [Inonotus] weirii was developed to assess potential control strategies in managed Pseudotsuga menziesii stands. The model mimicked key processes in disease initiation and development quantified as functions of time and space. These processes were horizontal and vertical tree root distribution, root contact with inoculum and among root systems, spread of mycelium through root systems, root decay, reduction of diam. growth in infected trees, tree mortality and persistence of inoculum in roots of stumps and killed trees. The processes were expressed as mathematical functions which were integrated in a computer program to calculate spread of the disease and stand-growth loss and mortality. Data for quantification of functions were obtained by experiments and from the literature. Simulated control practices included infected stump removal, sanitation fellings and mixed planting of Douglas fir and resistant species. Accuracy of the model was tested by comparing calculated disease spread and mortality with the following data: (1) spread and damage in two 60-yr-old, 1-ha stands in Oregon, (2) results from a statistically based model for spread and damage that had performed satisfactorily, and (3) observed spread and damage behaviour in stands of different ages and growth rates. Results from the model compared favourably with all of the above situations.

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10.
Brand, D.G. 1986a. A competition index for predicting the vigour of planted Douglas-fir in southwestern British Columbia. Canadian-Journal-of-Forest-Research 16(1): 23-29.

Keywords:      planting operations

                        tree/stand health

                        growth

Abstract: As a method of quantifying brush competition, data from 124 planted Douglas firs, age 1-5 yr, were used to derive a competition index to predict changes in tree vigour measured as a relative production rate. The index, which includes measures of brush proximity, relative ht. and % ground cover, appears to act as a measure of light interception around the tree crown. Tree vigour was found to be largely a function of the age of the tree from planting and the competition index. Foliage-based measures of growth vigour were related more strongly to the index than measures of b.a. or ht. The index has potential for assessing interspecific competition problems on suitable sites. Caution must be used in extrapolating results outside Douglas fir plantations on moist rich sites in coastal BC.

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11.
Brand, D.G. 1986b. Competition-induced changes in developmental features of planted Douglas-fir in southwestern British Columbia. Canadian-Journal-of-Forest-Research 16(2): 191-196.

Keywords:      planting operations

                        tree morphology

                        tree physiology

                        growth

Abstract: From measurements in 1- to 5-yr-old plantations, developmental characteristics of Douglas fir were tested against a competition index based on measures of the brush canopy surrounding individual trees. The most promising characteristics for assessing competition were specific leaf area, the allometric relationship of ht. to b.a. and bud production on nodal shoots. Measures of foliar N and leaf internode length were less well correlated with the competition index. Comparing these results with those of laboratory studies indicated that, on the study sites, brush competition effects on planted trees are expressed through adaptation to reduced light intensity. Developmental variables relating to moisture and nutritional status were not as strongly related to the competition index. This may reflect reduced tree demand or secondary brush canopy effects.

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12.
Brandeis, T.J., M. Newton and E.C. Cole. 2001. Underplanted conifer seedling survival and growth in thinned Douglas-fir stands. Canadian-Journal-of-Forest-Research 31(2): 302-312.

Keywords:      planting operations

                        thinning

                        commercial thinning

                        site preparation

                        chemical preparation

                        release treatments

                        chemical release

                        growth

                        tree/stand health

                        regeneration

Abstract: In a multilevel study conducted at the Oregon State University's McDonald-Dunn Research Forest, Oregon, USA, to determine limits to underplanted conifer seedling growth, Douglas-fir (Pseudotsuga menziesii), grand fir (Abies grandis), western redcedar (Thuja plicata) and western hemlock (Tsuga heterophylla) seedlings were planted in January 1993 beneath second-growth Douglas-fir stands that had been thinned in 1992 to basal areas ranging from 16 to 31 m2/ha. Understorey vegetation was treated with a broadcast herbicide (glyphosate + imazapyr) application prior to thinning, a directed release herbicide (glyphosate, plus triclopyr for tolerant woody stems) application 2 years later, or no treatment beyond harvest disturbance. Residual overstorey density was negatively correlated with percent survival for all four species. Broadcast herbicide application improved survival of grand fir and western hemlock. Western redcedar, grand fir and western hemlock stem volumes were inversely related to overstorey tree density and this effect increased over time. There was a strong indication that this was also the case for Douglas-fir. Reduction of competing understorey vegetation resulted in larger fourth-year stem volumes in grand fir and western hemlock.

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13.
Brandeis, T.J., M. Newton and E.C. Cole. 2002. Biotic injuries on conifer seedlings planted in forest understory environments. New Forests 24:1-14.

Keywords:      planting operations

                        site preparation

                        chemical preparation

                        release treatments

                        chemical release

                        thinning

                        tree/stand protection

                        growth

                        tree/stand health

Abstract: The effects of partial overstorey retention, understorey vegetation management, and protective Vexar(R) tubing on the frequency and severity of biotic injuries in a two-storied stand underplanted with western redcedar (Thuja plicata), Douglas-fir (Pseudotsuga menziesii), grand fir (Abies grandis), and western hemlock (Tsuga heterophylla) were investigated. The most prevalent source of damage was browsing by black-tailed deer (Odocoileus hemionis columbiana); deer browsed over 74% of Douglas-fir and over 36% of western redcedar seedlings one or more times over the four years of this study. Neither the spatial pattern of thinning (even or uneven) nor the density of residual overstorey affected browsing frequency. Spraying subplots may have slightly increased browsing frequency, but the resulting reduction of the adjacent understorey vegetation increased the volume of all seedlings by 13%, whether or not they were browsed. Vexar(R) tubing did not substantially affect seedling survival, browsing damage frequency, or fourth-year volume. Greater levels of overstorey retention reduced frequency of second flushing. Chafing by deer and girdling by rodents and other small mammals began once seedlings surpassed 1 m in height. Essentially all grand fir seedlings exhibited a foliar fungus infection.

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14.
Cameron, I.R. 1988. An evaluation of the density management diagram for coastal Douglas-fir. BC Ministry of Forests FRDA-Report 024. vi + 17 p.

Keywords:      planting operations

                        growth

                        yield

Abstract: Yield predictions based on Drew and Flewelling's (1979) density management diagram for coastal Douglas fir (Pseudotsuga menziesii) were compared with remeasured plot data from British Columbia. The diagram overestimated standing volume by 64% and mean diameter by 24% on average. Volumes predicted for plantations established at initial densities between 300 and 3000 trees per hectare exceeded nearly all the plots in the database. Because of the unattainable production targets set by the diagram, strategic plans based on its predictions would be seriously in error. Consequently, the diagram cannot be used in its current form as a silvicultural planning tool in stands of Douglas fir on the coast of British Columbia.

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15.
Carter, R.E., I.M. Miller and K. Klinka. 1986. Relationships between growth form and stand density in immature Douglas-fir. Forestry-Chronicle 62(5): 440-445.

Keywords:             planting operations
                               wood quality
                               tree morphology
                               growth

Abstract: Spacing was found to affect stem and crown characteristics and branching at whorls 6-10 in 27-yr-old Douglas fir established at 3 spacings (1.8x1.8, 3.6x3.6 and 4.6x4.6 m) near Haney, British Columbia. Ht., d.b.h. and stem diam. at the base of the live crown increased significantly with spacing, while age and ht. at the base of the live crown decreased. Increased spacing resulted in significantly greater branch diam. at whorls 6-10 and, in some whorls, an increase in branch number. Swelling of the stem at branch whorls, the number of Lammas whorls, knottiness ratio and a subjective index of stem form (where higher index indicated poorer form) increased with spacing. The selection of initial spacing is discussed and it is concluded that intensive management practices, e.g. thinning and pruning, will be necessary in all regimes if clear wood is to be produced over short rotations.

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16.
Childs, S.W. and L.E. Flint. 1987. Effect of shadecards, shelterwoods, and clearcuts on temperature and moisture environments. Forest-Ecology-and-Management 18(3): 205-217.

Keywords:      planting operations

                        tree/stand health

                        soil properties

                        tree physiology

                        tree phenology

Abstract: A comparison was made of two common techniques used to improve seedling survival on hot, dry reforestation sites. Adjacent shelterwood and clearcut sites in SW Oregon, USA, planted with 2+0 Douglas fir, were located and instrumented to compare temp. and moisture. In addition, cardboard shadecards were placed beside half of the seedlings studied. Seasonal measurements or observations of soil moisture, soil temp., solar radiation, air temp., stomatal diffusion resistance, seedling phenology and survival provided the basis for comparisons. Shelterwoods and shadecards improved seedling survival in relation to the clearcut. Both treatments affected soil temp. but the nature of the effects was different. The shelterwood canopy reduced solar radiation incident at the soil surface and caused cooler soil temp. throughout the soil profile. Shadecards reduced soil temp. only to a depth of 20 mm. Both treatments reduced the duration of periods of high soil temp. Shelterwood treatment delayed seasonal water loss and reduced seedling water stress as measured by stomatal resistance. Shadecards did not significantly affect seedling stomatal resistance. Differences in seedling survival caused by shadecards and shelterwoods are apparently due to different influences on the seedling microclimate. Shelterwood causes a large reduction in soil temp. as well as decreased seedling water stress. Shadecards modify the soil temp. less extensively and so have less effect on seedling survival.

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17.
Childs, S.W., H.R. Holbo and E.L. Miller. 1985. Shadecard and shelterwood modification of the soil temperature environment. Soil-Science-Society-of-America-Journal 49(4): 1018-1023.

Keywords:      planting operations

                        soil properties

Abstract: A study was conducted on steep, south-facing slopes in southwest Oregon to assess the effect of two common reforestation practices on the soil thermal environment. Three clearcut sites and three shelterwood sites were instrumented to measure soil temperature at five depths in the vicinity of shaded and unshaded Douglas-fir seedlings. Since the soils studied were skeletal, heat capacities of both fine and coarse soil fractions were determined for each site. These data were used to estimate soil heat fluxes. Shelterwoods decrease soil temperatures approximately 6 K when compared with clearcuts. This result holds at both 20- and 320-mm depths. Shelterwoods also decrease the depth of diurnal heating and decrease maximum hourly heat loss and gain values by 73 and 80 W/msuperscript 2, respectively. Shadecards, cardboard rectangles placed to the southwest of seedlings, generally have little effect on the soil temperature regime of skeletal soils but are effective in reducing daily heat flux. The dominant shadecard effect is a decrease in average daytime heat flux by 22 W/msuperscript 2, but shadecards also decrease average nighttime fluxes. Shelterwoods ameliorate seasonal soil temperature conditions significantly and may be an appropriate technique in situations where cumulative soil heating limits reforestation success. Shadecards should be useful in situations where heat stress events of only a few days are a problem. Over a season, shadecards exert little control, and their influence on stress is limited. Since all soils studied had high heat capacities due to large rock fragment content the conclusions of this study may be limited to soils with large soil heat capacity.

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18.
Colinas, C., R. Molina, J. Trappe and D. Perry. 1994a. Ectomycorrhizas and rhizosphere microorganisms of seedlings of Pseudotsuga menziesii (Mirb.) Franco planted on a degraded site and inoculated with forest soils pretreated with selective biocides. New-Phytologist 127(3): 529-537.

Keywords:      planting operations

                        fertilization

                        soil properties

                        mycorrhizal response

Abstract: Inoculation of planting holes with small amounts of soil from a mature forest or a plantation can improve formation of ectomycorrhizas on Pseudotsuga menziesii seedlings in degraded clearcuts in southwestern Oregon. To determine the component(s) of transferred soil responsible for increased ectomycorrhiza formation, soil from a clearcut, a mature forest and a plantation was treated with one of the following: (1) fertilizer to test for the effect of nutrients, (2) dimethoate and carbofuran to test for the effect on microarthropods or nematodes, (3) fumagillin to test for the effect on protozoa, (4) captan to test for the effect on fungi, (5) penicillin and oxytetracycline to test for the effect on bacteria, (6) pasteurization to test for the effect of active forms of organisms, (7) Tyndallization to test for the effect of resting forms of organisms, or (8) water as a control. The effect was studied of inoculation with soil subjected to these treatments on number and types of ectomycorrhizas, on length of active mycelium, and on number of active bacteria in the rhizosphere. Inoculation with untreated forest or plantation soils increased the number of ectomycorrhizas but did not change the mycorrhizal types present. Most agents had different effects in different soils. Inoculation with pasteurized and Tyndallized clearcut and plantation soils increased the number of Rhizopogon- and Thelephora-type ectomycorrhizas and decreased the number of active bacteria, as did untreated forest soil. It is hypothesized that the role of the soil transfer is to provide a rhizosphere environment free from a deleterious organism present in the clearcut. In this environment, beneficial organisms present in the clearcut or brought in with the seedling from the nursery can proliferate.

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19.
Colinas, C., D. Perry, R. Molina and M. Amaranthus. 1994b. Survival and growth of Pseudotsuga menziesii seedlings inoculated with biocide-treated soils at planting in a degraded clearcut. Canadian-Journal-of-Forest-Research 24(8): 1741-1749.

Keywords:      planting operations

                        fertilization

                        growth

                        tree/stand health

                        tree morphology

Abstract: To determine the factors of transfer soils responsible for increased seedling survival and growth, planting holes, at a site in SW Oregon, were inoculated with forest, plantation, and clear-cut soils subjected to one of 8 treatments: (i) treated with fertilizer to test for effects of nutrients; treated with biocides to test for effects of (ii) grazers (microarthropods or nematodes), (iii) protozoa, (iv) fungi, or (v) bacteria; (vi) pasteurized; (vii) Tyndallized; or (viii) untreated. Douglas fir (Pseudotsuga menziesii) seedlings were planted in June 1990 and seedling growth and survival was assessed in December 1990. Survival was increased by inoculation with untreated plantation soils, but not if they were fertilized or treated with dimethoate + carbofuran (grazercide), fumagillin (protozoacide), or oxytetracycline + penicillin (bactericide). Addition of untreated forest soil did not increase survival. For all soils, survival was increased by captan (fungicide), pasteurization and Tyndallization. Untreated plantation and forest soil transfers increased dry weights whereas neither did when treated with dimethoate + carbofuran. Dry weights of seedlings given clear-cut soil were increased by fertilization, pasteurization and Tyndallization of the soil; the latter two treatments also increased the number of short roots. It is hypothesized that stimulation of seedling growth by soil transfers was related to an increased rate of nutrient mineralization due to microbivorous soil animals contained within the transfer soils. Soil transfers may have enhanced seedling survival by at least two mechanisms: (i) by providing a safe site for beneficial rhizosphere organisms to proliferate, free from competing organisms that have proliferated in the clear-cut soil; and (ii) through volatile organic compounds, especially ethylene, that stimulated seedling root growth.

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20.
Comeau, P. and D. Sachs. 1992. Simulation of the consequences of red alder management on the growth of Douglas-fir using FORCYTE-11. B.C. Ministry of Forests FRDA Report 187. 45 p.

Keywords:      planting operations

                        yield

                        computer modeling

Abstract: The ecosystem model FORCYTE-11 was used to investigate the effects of initial red alder (Alnus glutinosa) density on yields of Pseudotsuga menziesii and alder over a 80-year rotation, the effects of delayed planting of red alder on stand yields, and the effects of 5 management strategies on the total yield over a 240-year period.

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21.
Copes, D.L. and M. Bordelon. 1994. Effects of tree spacing and height reduction on cone production in two Douglas-fir seed orchards. Western-Journal-of-Applied-Forestry 9(1): 5-7.

Keywords:      seed orchard management

                        planting operations

                        reproduction

Abstract: Two treatments involving tree spacings (12 and 24 ft) within rows spaced 24 ft apart and height control (topped or not topped at 20 ft) were evaluated in 17- and 22-yr-old coast Douglas fir (Pseudotsuga menziesii var. menziesii) seed orchards in Oregon. Treatment differences for cone production were not significant because of the large tree-to-tree variation in cone production and, at one orchard, the confounding of location with treatment. Cone production in 1990 averaged slightly more than 2 bushels per tree (range 0 to 11.7 bushels). Average production per 96 linear ft of orchard row was 14.8 bushels.

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22.
Curtis, R.O., G.W. Clendenen and D.J. DeMars. 1981. A new stand simulator for coast Douglas-fir: DFSIM user's guide. Pacific-Northwest-Forest-and-Range-Experiment-Station,-USDA-Forest-Service General-Technical-Report PNW-GTR-128. ii + 79 p.

Keywords:      planting operations

                        thinning

                        precommercial thinning

                        commercial thinning

                        fertilization

                        yield

                        computer modeling

Abstract: A description of a computer program, written in FORTRAN IV, for simulating managed stands. The program has been developed from remeasured plot data contributed by many organizations in the Pacific Northwest USA. It can produce yield tables which include estimates of effects of initial spacing, precommercial and commercial thinning and addition of N fertilizer. Topics discussed include program limitation and potential for further development. Appendices include operating instructions and notes on testing. The program is available from the authors on request.

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23.
DeYoe, D.R., H.R. Holbo and K. Waddell. 1986. Seedling protection from heat stress between lifting and planting. Western-Journal-of-Applied-Forestry 1(4): 124-126.

Keywords:      nursery operations

                        planting operations

                        tree/stand protection

                        tree/stand health

Abstract: Seven protective treatments were evaluated for preventing overheating of 2+0 Douglas fir seedlings in Kraft paper bags. Trials were conducted in May 1982 at Corvallis, Oregon on 3 clear days with max. air temp. of 78 degrees F and a hazy day with max. temp. 66 degrees . Seedlings were returned to cold storage (35 degrees ) overnight. Seedling temp. differed significantly between treatments. Unprotected seedlings (paper bag only) in full sun reached 89 degrees after 7 h. Green canvas caused increased heating rates and higher temp. (104 degrees after 7 h). A white sheet and a crinkled foil wrap performed no better than a paper bag alone. Canvas painted off-white reduced max. temp. to 80 degrees . Heavy shading (2% of full sun) and Mylar with white surface towards the sun were the most effective materials for preventing overheating (max. temp. 59-60 degrees ). Mylar with the silver surface facing the sun was less effective (max. temp. 71 degrees ).

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24.
Donegan, K.K., L.S. Watrud, R.J. Seidler, S.P. Maggard, T. Shiroyama, L.A. Porteous and G. DiGiovanni. 2001. Soil and litter organisms in Pacific Northwest forests under different management practices. Applied Soil Ecology 18:159-175.

Keywords:      planting operations

                        site preparation

                        prescribed fire

                        soil properties

Abstract: Soil and litter organisms were monitored for their response to different forest management practices. Litter and soil cores (0-10, 10-20 cm) were collected at approximately 8-week intervals over a 19-month period from a low elevation 110-140-year-old Douglas fir (Pseudotsuga menziesii) forest and adjacent 8-year-old clearcut in the Oregon Cascades, and from a high elevation 200-250-year-old Douglas fir forest and adjacent 5-year-old clearcut in the McKenzie River drainage. The low elevation clearcut had been broadcast burned and replanted with Douglas fir trees and a grass (Dactylis and Lolium)-legume (Trifolium and Lotus) mixture whereas the high elevation clearcut was not burned, large woody debris was left, and it was replanted with Douglas fir, Noble fir [Abies procera], Grand fir [Abies grandis], and western white pine [Pinus monticola]. The litter and soil cores were analysed for types of microarthropods and numbers of nematodes, fungi, culturable, aerobic bacteria, spore-forming bacteria, and chitin-degrading bacteria. Microbial community metabolic profiles, using the Biolog method, were also generated for the 0-10 cm soil samples. Populations of Pseudomonas spp. were analysed in the litter and soil samples using 16S rDNA fingerprints. Plant surveys were conducted to identify potential relationships of soil organisms to plant community composition. At both elevational field sites, there were significantly (P < 0.05) higher levels of nematodes and microarthropods in litter and soil in forest plots than in clear felled plots. Bacterial and fungal populations were also significantly higher in litter in forest plots than in clear felled plots at the high elevation site. In the litter and soil at the low elevation site and the soil at the high elevation site, however, microbial levels were higher in clear felled plots than in forest plots. The Pseudomonas spp. populations and the microbial community metabolic profiles in the 0-10 cm soil differed significantly between the forest and clear felled plots at the low elevation site but not at the high elevation site. At both elevational field sites, the plant cover (%) and plant density were significantly higher in clear felled plots than in forest plots. These observed differences in the population size and composition of organisms between mature forests and both low management and high management clearcuts demonstrated the impacts forest management practices may have on the soil ecosystem.

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25.
Dunsworth, B.G. 1988. Douglas-fir fall root phenology: a bioassay approach to defining planting windows. In Proceedings: 10th North American Forest Biology Workshop, 'Physiology and genetics of reforestation', University of British Columbia, Vancouver, British Columbia, July 10-22, 1988. Eds. J. Worrall, J. Loo-Dinkins and D.P. Lester. pp. 295-307.

Keywords:      planting operations

                        growth

                        tree/stand health

                        tree phenology

Abstract: Natural seedling root phenology during the autumn and spring was observed in Douglas fir [Pseudotsuga menziesii] and western hemlock [Tsuga heterophylla] over several seasons on Vancouver Island, British Columbia. These observations were used to determine the soil climate conditions under which peak activity occurred. Mitotic indexing and total root tip counts were used to quantify root activity. Peak activity for both spring and autumn in both species could be reasonably bracketed by soil climate conditions of -1 bar soil tension and 4 degrees C. The hypothetical planting window defined by these soil climate conditions was then tested with a series of timing of planting studies for Douglas fir (autumn) and western hemlock (spring) containerized stock. The results indicated that although quality of planting stock and the season following planting are influential, a 10 to 15% survival and growth advantage can be gained by planting within the hypothetical window.

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26.
Fight, R.D., S. Johnston, D.G. Briggs, T.D. Fahey, N.A. Bolon and J.M. Cahill. 1995. How much timber quality can we afford in coast Douglas-fir stands? Western-Journal-of-Applied-Forestry 10(1): 12-16.

Keywords:      pruning

                        planting operations

                        wood quality

                        economics

Abstract: Once site and genetic stock are selected, management of stocking, rotation age, and pruning are the principal means available to foresters to affect wood quality and value in stands of coast Douglas fir (Pseudotsuga menziesii var. menziesii) in the Pacific Northwest. Financial evaluation was used to test whether or not improvements in wood quality and value by these means justify the cost of doing so. This analysis showed in general that improving quality through high levels of stocking or extending rotations were costly ways to improve wood quality while pruning was cost effective.

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27.
Grotta, A.T., B.L. Gartner and S.R. Radosevich. 2004. Influence of species proportion and timing of establishment on stem quality in mixed red alder-Douglas-fir plantations. Canadian-Journal-of-Forest-Research 34(4): 863-873.

Keywords:      planting operations

                        tree morphology

                        wood quality

Abstract: The relationships among stand structure, Douglas-fir (Pseudotsuga menziesii) branch characteristics, and red alder (Alnus rubra) stem form attributes were explored for 10- to 15-year-old trees growing in mixed Douglas-fir-red alder plantations in Oregon, USA. Treatments included a range of species proportions, and red alder was either planted simultaneously with Douglas-fir or after 5 years. Both replacement effects (total stand density held constant) and additive effects (stand density doubled) of competition were considered. When the two species were planted simultaneously and red alder proportion was low, red alder trees had low crown bases and much stem defect (lean, sweep, and multiple stems). Douglas-fir grew slowly when the two species were planted simultaneously. When red alder planting was delayed, species proportion did not affect red alder stem form, and height to the base of the Douglas-fir live crown decreased with increasing red alder proportion. Doubling Douglas-fir density increased the height to the base of the Douglas-fir live crown; however, doubling stand density by adding red alder did not affect Douglas-fir crown height. Douglas-fir lumber coming from mixed stands may be inferior because of the changes in knot characteristics associated with these different patterns of crown recession. In stands with a low proportion of red alder, red alder product recovery may be compromised because of the stem defects described above.

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28.
Haase, D.L., J.H. Batdorff and R. Rose. 1993. Effect of root form on 10-year survival and growth of planted Douglas-fir trees. Tree-Planters' Notes 44(2): 53-57.

Keywords:      planting operations

                        growth

                        tree/stand health

Abstract: Douglas fir seedlings (Pseudotsuga menziesii) were planted with three root-form treatments including C-roots ('correctly' planted controls), L-roots, and J-roots. After 10 years, there were no significant differences in outplanting performance between the three root-form treatments on a good site in western Oregon. The results are in agreement with those of other studies, which suggests that when no other confounding planting errors are present, deformed root systems play a less dramatic role in subsequent field performance than is generally thought. These results in no way imply that poor planting is acceptable.

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29.
Hahn, P.F. and A.J. Smith. 1983. Douglas-fir planting stock performance comparison after the third growing season. Tree-Planters' Notes 34(1): 33-39.

Keywords:      nursery operations

                        planting operations

                        growth

                        tree/stand health

Abstract: Three types of containerized (40, 75 or 125 cmsuperscript 3 containers) and bare rooted (2+1, 3+0 and plug-1 stock) seedlings were planted out in Oregon in Feb. 1979 on N. and S. facing slopes, clear felled in 1978. In general, containerized seedlings showed superior survival rates and greater height growth - particularly on the harsh S. slope, and lower reforestation costs. The 75-cmsuperscript 3 containerized seedlings are recommended, except for N. facing and wet coastal areas, where brush competition can be severe shortly after planting. In such areas, taller bare-rooted seedlings performed better.

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30.
Haight, R.G. 1993a. The economics of Douglas-fir and red alder management with stochastic price trends. Canadian-Journal-of-Forest-Research 23(8): 1695-1703.

Keywords:      planting operations

                        site preparation

                        prescribed fire

                        release treatments

                        chemical release

                        thinning

                        precommercial thinning

                        commercial thinning

                        tree/stand protection

                        economics

Abstract: A financial analysis of Douglas fir (Pseudotsuga menziesii) and red alder (Alnus rubra) management was conducted using yield projections from the Stand Projection Simulator for the Pacific Northwest region of the United Sates. The analysis included uncertainty in the price trends and stocking levels of both species following reforestation. Results from a case study in which Douglas fir price is likely to increase faster than red alder price show that (i) on more productive sites, greater regeneration investment is justified to increase the likelihood of Douglas fir establishment, (ii) on less productive sites, low-cost regeneration options that produce mixed-species stands have expected present values close to or greater than a high-cost Douglas fir regeneration effort, (iii) optimal precommercial removal of red alder depends on mid-rotation prices and regeneration success, and in many cases growing a mixed-species stand to maturity produces the highest economic return, and (iv) commercial thinning of Douglas fir increases the expected present value of the most intensive regeneration option by up to 10%. The low-cost regeneration options have relatively high expected returns because of low initial investments and the presence of two species that may have high values in the future. The sensitivity of these results to changes in the probability distributions of regeneration success and price trends is discussed.

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31.
Haight, R.G. 1993b. Technology change and the economics of silvicultural investment. Rocky-Mountain-Forest-and-Range-Experiment-Station,-USDA-Forest-Service General-Technical-Report RM-GTR-232. ii + 18 p.

Keywords:      planting operations

                        site preparation

                        prescribed fire

                        release treatments

                        chemical release

                        tree/stand protection

                        thinning

                        commercial thinning

                        precommercial thinning

                        yield

                        economics

Abstract: Financial analyses of intensive and low-cost reforestation options are conducted for loblolly pine (Pinus contorta) stands with broadleaved competition in the Southern USA, and Douglas fir with red alder (Pseudotsuga menziesii with Alnus rubra) in the Pacific Northwest. Results show that the expected present values (EPVs) of low-cost options that result in mixtures of conifers and broadleaves are superior in some situations to the EPVs of the intensive options.

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32.
Heilman, P. 1983. Effects of surface treatment and interplanting of shrub alder on rowth of Douglas-fir on coal spoils. Journal-of-Environmental-Quality 12(1): 109-113.

Keywords:      planting operations

                        site preparation

                        mechanical preparation

                        growth

                        tree physiology

                        soil properties

                        tree/stand health

Abstract: Annual growth of Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) planted on topsoiled spoils at a coal mine near Centralia, Wash., was monitored for the first 6 y after planting. Treatments were contour bedding, contour bedding plus interplanted Sitka alder (Alnus sinuata (Reg.) Rydb.), and unbedded control. The bedding significantly increased growth of Douglas-fir in all 5 y of the study. Total height growth after 5 y was 35% greater than control on the bedding only plots, and 43% greater on the bedding plus Sitka alder plots. Height growth of Douglas-fir in the mixed stand was significantly greater during the 2nd and 3rd y of the study, but after 5 yr, no significant difference was evident in total height between the mixed and pure Douglas-fir plots. Concentration of N in Douglas-fir foliage was significantly increased by bedding in the fifth but not in the fourth year. Interplanting with Sitka alder had no significant effect on N in Douglas-fir foliage. The top 0.3 m of soil in the ridged portion of the bedded area contained significantly less moisture over a summer than did the top 0.3 m of the unbedded soil. At deeper depths, however, soil moisture was not significantly affected by bedding. Wind damage caused by a severe storm that occurred after 5 y was very much greater on the unbedded plots (49% wind-thrown vs. 9 to 15% wind-thrown on the bedded plots) despite the smaller size of the trees on unbedded plots.

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33.
Helgerson, O.T. 1985. Survival and growth of planted Douglas-fir (Pseudotsuga menziesii (Mirb.) Franco) and ponderosa pine (Pinus ponderosa Dougl. ex Laws.) on a hot, dry site in southwest Oregon. Tree-Planters' Notes 36(4): 3-6.

Keywords:      nursery operations

                        planting operations

                        tree/stand health

                        growth

Abstract: One-yr-old containerized seedlings and 2-yr-old bare rooted seedlings of both species were planted in Feb. 1982 on a W.-facing 35% slope on Tin Pan Peak. The site receives <760 mm of precipitation annually. Weeds were controlled with herbicides applied before and after planting. Survival after 2 yr averaged 94% for all stock types; survival of bare rooted seedlings (98-99%) was significantly better than that of containerized seedlings (88-92%). Relative vol. growth was greater for pine than Douglas fir. After 2 yr, the 2+0 bare rooted pines were significantly larger than the 2+0 Douglas firs, despite a smaller starting size.

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34.
Helgerson, O.T. 1990a. Effects of alternate types of microsite shade on survival of planted Douglas-fir in southwest Oregon. New-Forests 3(4): 327-332.

Keywords:      planting operations

                        tree/stand protection

                        tree/stand health

                        growth

Abstract: Five-yr survival of 2+0 bare root Douglas fir (Pseudotsuga menziesii) seedlings was increased by 3 types of shading: cardboard shadecards placed S. or E. of seedlings; and bottomless styrofoam cups inverted around seedling base, on 2 S.-facing sites. On the drier site, seedlings survived well without shading (89% unshaded, 98% shaded), but on the wetter site, where seedlings were more stressed, shading was more beneficial (62% unshaded, 89% shaded). Shading did not affect growth. Seedlings grew more in 5 yr on the drier than the wetter site, possibly because of better handling and planting practices, less browsing by deer, and better weed control.

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35.
Helgerson, O.T., S.D. Tesch, S.D. Hobbs and D.H. McNabb. 1989. Survival and growth of ponderosa pine and Douglas-fir stocktypes on a dry low-elevation site in southwest Oregon. Western-Journal-of-Applied-Forestry 4(4): 124-128.

Keywords:      nursery operations

                        planting operations

                        growth

                        tree/stand health

Abstract: Two stocktypes (1+0 container-grown plugs and 2+0 nursery-grown bareroots) of ponderosa pine (Pinus ponderosa) and of Douglas fir (Pseudotsuga menziesii) were planted on a hot, droughty, low-altitude site near Medford, Oregon, which had burned in 1981. The main objective was to assess the potential for reforesting this type of site. After 5 growing seasons, bare rooted stock survived (98%) significantly better than plugs (89%); survival did not differ significantly by species. Douglas fir was taller than pine, pine was larger in diam., and the 2 species had approximately equal stem volumes. Bare rooted stock was consistently larger than plugs. Results show that these species and stocktypes can provide good reforestation after 5 yr on this type of site when seedlings are of good quality, are planted properly, and are given good weed control.

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36.
Helgerson, O.T., S.D. Tesch, S.D. Hobbs and D.H. McNabb. 1992. Effects of stocktype, shading, and species on reforestation of a droughty site in southwest Oregon. Northwest-Science 66(2): 57-61.

Keywords:      nursery operations

                        planting operations

                        tree/stand protection

                        tree/stand health

                        growth

Abstract: On hot, dry sites, shading may differentially increase survival of planted Douglas fir (Pseudotsuga menziesii) according to seedling size, and Douglas fir may differ from ponderosa pine (Pinus ponderosa) in early survival and growth. The survival and growth of Douglas fir seedlings (1+0 container-grown plugs and 2+0 bare-rooted seedlings, unshaded or shaded with cardboard shadecards at planting) and unshaded 2+0 bare-rooted ponderosa pine were compared on a droughty south facing clear felling in Oregon. The site was clear felled and burned in 1982 and the seedlings were planted in 1983. Shading did not significantly increase survival of plugs, possibly because of a wetter than normal first summer, nor did shading affect growth of either Douglas fir stocktype 5 yr after planting. Bare-rooted Douglas fir remained significantly larger than plugs, but relative growth rates for the initially smaller plugs were significantly greater for diameter and volume. Survival and growth of ponderosa pine tended to be better than those of Douglas fir. It was concluded that both species appeared to be suitable for reforestation after clear felling on sites subject to drought.

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37.
Hermann, R.K. and D.P. Lavender. 1999. Douglas-fir planted forests. New-Forests 17(1/3): 53-70.

Keywords:      genetic tree improvement

                        nursery operations