1. 1987. Impact of intensive forestry practices on net stand values in British Columbia. B.C. Ministry of Forests FRDA-Report 014. 109 p.

Keywords:      release treatments

                        fertilization

                        thinning

                        yield

                        economics

Abstract: Yield responses to major silvicultural treatments (regeneration method, brushing and weeding, spacing and thinning and fertilizer use) are analysed in relation to growth and yield theory, and their translation into operational use of treatments to increase merchantable vol. is considered. Data from coastal Douglas fir (Pseudotsuga menziesii), western hemlock (Tsuga heterophylla) and western hemlock/Abies amabilis stands and interior white spruce (Picea glauca), lodgepole pine (Pinus contorta) and wet belt Douglas fir stands are used to quantify the net present value of treatments in terms of improvement in net stand values and merchantable vol. Potentially viable treatment options are identified for each stand type present.

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2.
Anderson, H.W. and M. Hyatt. 1981. Feasibility of hand application of urea to forest land in western Washington. In Proceedings: Forest Fertilization Conference, University of Washington, Seattle, Washington, USA. Eds. S.P. Gessel, R.M. Kenady and W.A. Atkinson. pp. 205-208.

Keywords:      fertilization

                        economics

Abstract: A large scale pilot study to determine the potential of hand fertilizing forest stands in western Washington was undertaken by the Department of Natural Resources in 1976. Tests were conducted in two major stand types of Douglas-fir (juvenile stands with voids and openings and older open growh stands), where two hand spreading techniques (broadcast and individual tree fertilization) and two types of work crews (contract and DNR) were evaluated. During the study manhours of the various job-related activities were recorded and costs determined. A total of 634 acres was treated and costs ranged from $36.95 to $125.42 per acre and averaged $81.74 per acre (1976 dollars). Costs varied because of stand type which influenced the fertilizer rate and crew type which influenced the application cost. Based on units treated in this study and assuming similar conditions, estimates of hand fertilization costs for an operational fertilization project where stand conditions would allow for a reduced fertilizer rate (individual tree fertilization) would be approximately $52 per acre at the rate of 330 pounds of urea and $42 per acre at the rate of 220 pounds of urea. This compares to costs of $61 per acre for hand broadcast application at 440 pounds of urea and $56 per acre for aerial application at the same rate. Therefore, where stand conditions allow for a reduced rate of fertilizer, hand application could be a more inviting alternative to the normal procedure of aerial application.

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3.
Atkinson, W.A. 1981. Preliminary guidelines for fertilizing less than fully stocked stands and mixed species stands. In Conference Proceedings: Forest Fertilization Conference, University of Washington, Seattle, Washington, USA. Eds. S.P. Gessel, R.M. Kenady and W.A. Atkinson. pp. 59-61.

Keywords:      fertilization

                        economics

Abstract: Foresters engaged in selecting areas for operational fertilization rarely encounter the fully stocked pure-species Douglas-fir stands that are studied by researchers. Most often the question is one of establishing minimal standards for choosing stands to fertilize. This paper offers preliminary guidelines for fertilizing less than fully stocked stands and mixed-species stands. Results are presented by age and site, and consist of a table showing minimal basal area stocking required to earn a given interest rate.

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4.
Bailey, V.L., J.L. Smith and H. Bolton, Jr. 2002. Fungal-to-bacterial ratios in soils investigated for enhanced C sequestration. Soil-Biology-and-Biochemistry 34(7): 997-1007.

Keywords:      fertilization

                        soil properties

Abstract: Fungi and bacteria govern most of the transformations and ensuing long-term storage of organic C in soils. We assessed the relative contributions of these two groups of organisms to the microbial biomass and activity of soils from five different ecosystems with treatments hypothesized to enhance soil C sequestration: (1) desert (an elevation gradient allowed comparison of soil developed in a cooler, wetter climate with soil developed in a warmer, drier climate), (2) restored tallgrass prairie (land reverted to native prairie in 1979 and neighbouring land farmed to row crops for ~100 year), (3,4) two forest types (Douglas fir [Pseudotsuga menziesii] and loblolly pine [Pinus taeda], unfertilized control and N-fertilized plots), and (5) agricultural land (conventional- and no-till management systems). The selective inhibition technique, using captan (fungicide) and oxytetracycline hydrochloride (bactericide), was used to determine the activities (respiration) of fungi and bacteria in each of these soils and substrate-induced respiration was used to measure total active soil microbial biomass C. Phospholipid fatty acid analysis was used to determine the composition of the soil microbial biomass and determine if the activities and structure of the microbial communities were related. Differences in fungal-to-bacterial (F:B) activities between treatments at a site were greatest at the prairie sites. The restored prairie had the highest F:B (13.5) and high total C (49.9 g C kg-1 soil); neighbouring soil farmed to maize had an F:B of 0.85 and total C of 36.0 g C kg-1 soil. Within the pairs of study soils, those that were tilled had lower fungal activities and stored C than those that were managed to native or no-till systems. In all pairs of soils, soils that had higher absolute fungal activities also had more total soil C and when two extreme cases were removed fungal activity was correlated with total soil C (R2=0.85). Thus, in this small set of diverse soils, increased fungal activities, more than F:B ratios, were associated with increased soil C. Practices that involved invasive land management decreased fungal activity and stored soil C compared to similar soils that were less intrusively managed.

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5.
Ballard, T.M. 1984. A simple model for predicting stand volume growth response to fertilizer application. Canadian-Journal-of-Forest-Research 14(5): 661-665.

Keywords:      fertilization

                        growth

Abstract: The equation R=KTACZQ is used to predict cumulative stand volume growth response (R) to fertilizer application, where K is a constant and the last 5 (dimensionless) multipliers represent functions of time, amount of fertilizer applied, stand composition, stocking, and site quality, respectively. Site-specific input data requirements are stand composition expressed as percent responding species, stocking expressed as a percentage of normal stocking, and site index. The model was calibrated for Douglas-fir response to nitrogen using data from Washington and Oregon. A preliminary test of the model and its calibration compared predicted responses with estimates of actual response derived from some fertilizer trials on Vancouver Island. Soil drainage class and foliar analysis data may help in qualitatively inferring whether the model's site quality function leads to overestimation of response. Future development of the model may include development and calibration of a predictively better site quality function which quantitatively uses site water regime and stand nutrient data.

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6.
Ballard, T.M. and N. Majid. 1985. Use of pretreatment increment data in evaluating tree growth response to fertilization. Canadian-Journal-of-Forest-Research 15(1): 18-22.

Keywords:      fertilization

                        growth

Abstract: The use of pretreatment increment can lead to improved estimates of individual and average tree growth response to fertilizing, by helping to adjust for site as well as stand structure differences between fertilized and control areas. It has applications in research using either single-tree or plot fertilizing, and also in estimating responses to operational fertilizing. Equations for analysing increment response to fertilizing are presented and discussed and 2 particularly useful ones were evaluated by examining branch length increment data from foliar spray application of iron and copper to Pinus contorta and of nitrogen and iron to Pseudotsuga menziesii, in field trials in British Columbia.

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7.
Barclay, H., H. Brix and C.R. Layton. 1982. Fertilization and thinning effects on a Douglas-fir ecosystem at Shawnigan Lake: 9 year growth response. Pacific-Forestry-Centre, Canadian-Forest-Service Information-Report BC-X-238. 35 p.

Keywords:      fertilization

                        thinning

                        growth

Abstract: Further results are given for a trial established in 1970 in a 24-yr-old stand in British Columbia, last measured 6 yr after treatment. Volume increments over 9 yr for heavy thinning alone, heavy fertilizer (urea) treatment alone, or both together, were 46%, 75% and 120%, respectively.

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8.
Barclay, H.J. and H. Brix. 1984. Effects of urea and ammonium nitrate fertilizer on growth of a young thinned and unthinned Douglas-fir stand. Canadian-Journal-of-Forest-Research 14(6): 952-955.

Keywords:      fertilization

                        thinning

                        growth

                        tree physiology

                        tree/stand health

Abstract: The effects were studied of 2 sources of nitrogen fertilizer applied at rates of 224 and 448 kg/ha N on growth of thinned and unthinned plots established in 1970 in a 24-yr-old stand on southern Vancouver Is., British Columbia. Ammonium nitrate yielded higher growth of diam. and vol. than urea over a 9-yr period, particularly with thinning. Ht. growth was not affected by nitrogen source. The efficiency of nitrogen fertilizing in terms of stem vol. response per kilogram of nitrogen applied was greatest with ammonium nitrate in thinned plots. Tree mortality increased substantially with fertilizing for both sources, and decreased markedly with thinning.

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9.
Barclay, H.J. and H. Brix. 1985a. Effects of high levels of fertilization with urea on growth of thinned and unthinned Douglas-fir stands. Canadian-Journal-of-Forest-Research 15(4): 730-733.

Keywords:      fertilization

                        thinning

                        growth

                        tree physiology

                        tree/stand health

Abstract: Diameter height and volume growth were documented for 9 yr after thinning and fertilizing in a 24-yr-old stand on a poor site on southern Vancouver Is. The treatments involved 3 thinning treatments (0, 1/3, and 2/3 b.a. removed) and 6 fertilizer treatments (0-1344 kg/ha N) with urea. Increments for both diameter and gross volume increased with the rate of fertilizer application and responses were still apparent 9 yr after treatment. For unthinned plots, the 9-yr volume growth responses were 30, 50, and 80% with fertilizer rates of 224, 448, and 896 kg/ha N, respectively. The efficiency of fertilizer use, measured as stem volume response per unit of nitrogen applied, decreased with rate of fertilizer application, but this result may change over a longer response period. There was a positive interaction between fertilizing and thinning such that high amounts of both mutually enhanced growth. Mortality increased with fertilizing, but only noticeably in unthinned plots.

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10.
Barclay, H.J. and H. Brix. 1985b. Fertilization and thinning effects on a Douglas-fir ecosystem at Shawnigan Lake: 12-year growth response. Pacific-Forestry-Centre, Canadian-Forest-Service Information-Report BC-X-271. 34 p.

Keywords:      fertilization

                        thinning

                        growth

Abstract: [See FA 45, 2316] Further results are given from the study established in 1970 in a 24-yr-old stand in British Columbia. Fertilizers (urea) and thinning both increased vol. increments over 12 yr. Refertilization 9 yr after initial treatment has produced substantial increases in vol. increment.

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11.
Barclay, H.J. and C.R. Layton. 1990. Growth and mortality in managed Douglas fir: relation to a competition index. Forest-Ecology-and-Management 36(2-4): 187-204.

Keywords:      fertilization

                        thinning

                        growth

                        tree/stand health

Abstract: Twelve-year increments of diameter at breast height (DBH), height, and volume in thinned and fertilized 45-year-old Douglas fir (Pseudotsuga menziesii) stands on Vancouver Island, Canada, were related (by regression) to degree of thinning, amount of fertilizer (3x3 factorial), initial DBH, and a competitive stress index (CSI). The ability of the CSI to predict growth after treatment was examined. Causes of tree death, and CSI data, are presented, and the relationship between them discussed. The CSI was found to be only moderately good at predicting Douglas fir growth and mortality: initial DBH provided a better predictor. Most mortality in unthinned plots resulted from suppression, and correlated reasonably well with CSI; mortality in thinned plots was not correlated with CSI, and resulted principally from snow damage. Tree height variability generally became less over the 12 years following treatment, which is more consistent with two-sided than one-sided competition predictions, a result which is contrasted to that of many other species.

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12.
Barclay, H.J., P.C. Pang and D.F.W. Pollard. 1986. Aboveground biomass distribution within trees and stands in thinned and fertilized Douglas-fir. Canadian-Journal-of-Forest-Research 16(3): 438-442.

Keywords:      fertilization

                        thinning

                        carbon allocation

Abstract: Nine years after heavy thinning and fertilization with urea, 34-yr-old Douglas firs at Shawnigan Lake (British Columbia) were destructively sampled. Dry wt. of seven aboveground components (wood, bark, dead branches, new or old foliage, new twigs and live branches) were determined and regression equations from d.b.h. were developed. Differences among treatments were shown for all biomass components and for the proportion of the total biomass allocated to each component. Thinning reduced the proportion of wood, bark and dead branches while increasing the proportion of foliage and live branches. Fertilization increased the proportion of branches but had negligible effects on the proportions of other components.

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13.
Barclay, H.J. and J.A. Trofymow. 2000. Relationship of readings from the LI-COR canopy analyzer to total one-sided leaf area index and stand structure in immature Douglas-fir. Forest-Ecology-and-Management 132(2/3): 121-126.

Keywords:      fertilization

                        thinning

                        tree morphology

Abstract: Estimation of leaf area is important in predicting potential growth. This estimation is often done by means of a photometer, such as the LI-COR plant canopy analyser, but such instruments generally give biased estimates. Consequently, conversion factors are required to convert output from the photometer to the actual leaf area index (LAI). Foliar biomass was estimated in a 52-year-old Douglas fir (Pseudotsuga menziesii) stand at Shawnigan in British Columbia, Canada, which had been treated with 3 levels of thinning and 3 levels of fertilizer 28 years previously. The 4 treatment extremes (no thinning or fertilizing, no thinning and heavy fertilizing (448 kg N/ha as urea), heavy thinning (to 1/3 basal area) and no fertilizing, and heavy thinning and heavy fertilizing) were sampled for foliage. Projected leaf areas were calculated from these biomass samples using specific leaf areas derived in a previous biomass sampling. Total one-sided leaf area was then computed by dividing the projected leaf areas by 0.9 to allow for lateral leaf curvature. In addition, LAI-2000 readings were taken in the same plots and then factors were derived to convert LAI-2000 readings to total one-sided LAI. These conversion factors were found to vary strongly with quadratic mean diameter, stand density, mean diameter at breast height, mean height and stand basal area. The effect of live crown height was minimal. These should allow the prediction of the conversion factor based on 2 or 3 of these highly correlated factors.

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14.
Bare, B.B. 1981. Tax effects of fertilization. In Proceedings: Forest Fertilization Conference, University of Washington, Seattle, Washington, USA. Eds. S.P. Gessel, R.M. Kenady and W.A. Atkinson. pp. 238-242.

Keywords:      fertilization

                        economics

Abstract: This paper describes the sensitivity of investment performance when fertilzization activities are treated as expensed, capitalized, or amortized expenditures. Current Internal Revenue Service policy favors capitalization, but pending revenue rulings are expected to recommend amortization. The impact of these three alternatives on after-tax cash flows is illustrated by a numerical example for the Douglas-fir zone of the USA.

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15.
Binkley, D. and P. Reid. 1984. Long-term responses of stem growth and leaf area to thinning and fertilization in a Douglas-fir plantation. Canadian-Journal-of-Forest-Research 14(5): 656-660.

Keywords:      fertilization

                        thinning

                        tree morphology

                        carbon allocation

                        growth

Abstract: Replicated thinning and nitrogen fertilization plots in a 53-year-old plantation in Washington State were examined for repsonses in stem growth, leaf area, and stem growth per unit leaf area. Although measurements occurred 20-30 yr after plot installation, substantial effects from the various treatments were still present. Thinning reduced leaf area of the stands but increased stem growth per unit leaf area, resulting in little difference in stem growth per ha over the 5-yr measurement period (1977-81). Fertilization increased both stand leaf area and stem growth per unit leaf area, and more than doubled 5-yr stem growth per ha. Consideration of the role of leaf area and stem growth per unit leaf area in determining stand treatment responses may account for much of the variation found among replicates of treatments or between studies on different sites.

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16.
Binkley, D. and P. Reid. 1985. Long-term increase of nitrogen availability from fertilization of Douglas-fir. Canadian-Journal-of-Forest-Research 15(4): 723-724.

Keywords:      fertilization

                        growth

                        tree physiology

                        soil properties

Abstract: [See FA 44, 4708; 46, 1837] Most Douglas-fir stands respond to nitrogen fertilizing by increasing stem growth for less than 8 yr, but one plantation at the United States Forest Service Wind River Experimental Forest in Washington State has responded for over 15 yr. In this study nitrogen concn. of foliage and fresh litter were shown to be higher in the fertilized plots (470 kg/ha N) 18 yr after fertilizing. Retranslocation of N from senescent needles was not affected and stem growth per unit N in the canopy was similar between unfertilized and fertilized plots. An index of soil N availability in the fertilized plots was twice that of unfertilized plots. The higher stem growth, leaf area, and stem growth per unit leaf area demonstrated in an earlier study appeared to be related to a sustained increase in soil N availability rather than increased N-use efficiency. An examination of soil N transformation processes is needed to complete the explanation of the unusually prolonged fertilizer response in these plots.

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17.
Blake, J., S.R. Webster and S.P. Gessel. 1988a. Soil sulfate-sulfur and growth responses of nitrogen-fertilized Douglas-fir to sulfur. Soil-Science-Society-of-America-Journal 52(4): 1141-1147.

Keywords:      fertilization

                        soil properties

                        growth

Abstract: Two studies were conducted to determine the growth response of N-fertilized Douglas fir [Pseudotsuga menziesii (Mirb.) Franco] to S supplements. The relationship between response and soil SO4-S extracted with Morgan's solution, 1.22 M NaOAc + 0.53 M HOAc (pH 4.8), was used to establish critical levels for S. Douglas fir seedlings were grown in the greenhouse in the surface mineral layer (0 to 0.15 m) of 20 forest soils from western Washington and Oregon. On the average, significant increases in total dry weight (17.5%), stem diameter (10.1%), and height (6.9%) occurred when soils were fertilized with N and S in comparison to N alone. Using the Cate-Nelson procedure, growth responses to N and S were most likely to occur when soil SO4-S was below 14 mg S kg-1. Twenty eight installations were established in the field containing five treatments, three rates of N as urea, and one plot of 336 kg N ha-1 with P, K, Ca, and S. Differences in percent basal area growth between N alone and N with P, K, Ca, and S were significantly related to soil SO4-S. Over the initial 5-yr period, response over N alone was improved by 74% when soil SO4-S was <20 mg S kg-1. When the N with P, K, Ca, and S plots were retreated with only N and S, response over the next 3 yr was more than doubled compared with N alone. Identification of S responsive stands was improved by the inclusion of stand age weighted subsoil SO4-S concentrations.

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18.
Blake, J.I., H.N. Chappell, W.S. Bennett, S.R. Webster and S.P. Gessel. 1990. Douglas fir growth and foliar nutrient responses to nitrogen and sulfur fertilization. Soil-Science-Society-of-America-Journal 54(1): 257-262.

Keywords:      fertilization

                        tree physiology

                        growth

Abstract: Nitrogen-fertilizer response in conifer stands of the Pacific Northwest has been related to soil and foliar S, and growth has sometimes been enhanced by the addition of S. Five stands of Douglas fir (Pseudotsuga menziesii) in Washington or Oregon, with low to moderate quantities of sulfate in the mineral soil horizon, were treated with N (urea) alone or with S (as ammonium sulfate). The results indicated that levels of soil sulfate did not provide local or site-specific predictions of the magnitude of the gain from applying N with S. The observed treatment effects were highly variable. Foliar N concentrations in the N plus S treatment were generally higher than in the N treatment. Little change in foliar S content occurred in the N plus S plot. Periodic annual growth response to N over the study period was inversely related to site index and directly related to foliar N content.

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19.
Bodner, J. 1984. Effect of thinning and fertilization on wood properties and intra-ring characteristics in young Douglas-fir. Holzforschung-und-Holzverwertung 36(1): 5-11.

Keywords:      fertilization

                        thinning

                        wood quality

Abstract: Studies were made on samples from a total of 21 trees (felled in 1982) from 42-yr-old control and thinned/[N] fertilizer-treated stands near Sweet Home, Oregon, and a 48-yr-old thinned stand near Corvallis. Wood properties, studied between and within treatments, included ring density (analysed by X-ray densitometry), earlywood and latewood density, min. earlywood density, max. latewood density, and ring width. There was n.s.d. in av. wood density between treatments. There were significant between-treatment differences in MOE and MOR. Heavy thinning (during the juvenile wood formation phase) reduced latewood fibre length by 26.5%. Min. earlywood density and max. latewood density were the most important components of ring density.

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20.
Briggs, D.G., F. Mecifi and W.R. Smith. 1986. Effect of sludge on wood properties: a conceptual review with results from a sixty-year-old Douglas-fir stand. In The forest alternative for treatment and utilization of municipal and industrial wastes. Ed. D.W. Cole, C.L. Henry, and W.L. Nutter. Seattle, Washington, USA: University of Washington Press. pp. 246-257.

Keywords:      fertilization

                        wood quality

Abstract: Expected changes in wood properties due to cultural practices are reviewed. The hormone theory and published studies on the effects of thinnings and fertilizers are used to provide a basis for hypotheses describing the effects of sludge treatments on wood properties. Notes are given on the effect of municipal sludge on relative density, tracheid characteristics and strength properties of Douglas fir.

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21.
Brix, H. 1984. Effects of thinning and nitrogen fertilization on growth of Douglas-fir: relative contribution of foliage quantity and efficiency. Canadian-Journal-of-Forest-Research 13(1): 167-175.

Keywords:      fertilization

                        thinning

                        growth

                        carbon allocation

                        tree morphology

Abstract: [See FA 43, 1948, 3839] On Vancouver Island, aboveground biomass and annual production over 7 yr was studied in relation to thinning and nitrogen fertilization at 24 yr old. Biomass yield of both treatments increased during the first 3-4 yr then decreased for fertilization but not with thinning. Treatments doubled biomass production of individual trees over the study period when applied separately and quadrupled it when combined. Annual biomass production per unit of foliage (E) increased during the first 3-4 yr, but was at or below control level after 7 yr. E accounted for 20, 37, and 27% of the stemwood dry matter response to thinning, fertilization and the combined treatments, respectively; the remainder was attributed to an increase in foliage biomass. Thinning, but not fertilization, influenced distribution of radial growth along the stem, increasing growth only below the top one-third of the stem. This pattern was related to crown development.

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22.
Brix, H. 1993. Fertilization and thinning effect on a Douglas-fir ecosystem at Shawnigan Lake: a synthesis of project results. B.C. Ministry of Forests FRDA-Report 196. X + 64 p.

Keywords:      fertilization

                        thinning

                        growth

                        tree morphology

                        tree/stand health

                        carbon allocation

                        wood quality

                        tree physiology

                        photosynthesis

                        economics

Abstract: Treatments were initiated in 1970-71 in a 24-year-old Douglas fir (Pseudotsuga menziesii) near Shawnigan Lake, Vancouver Island, British Columbia, to determine the effects of 3 intensities of thinning (removing none, one-third and two-thirds of basal area) and 3 levels of urea fertilizer (0, 224 and 448 kg N/ha) on the growth and biology of the trees. Subsidiary experiments were established during 1972-87 to examine the effects of high doses of urea (672-1344 kg N/ha), ammonium nitrate as an N source instead of urea, understorey response to thinning and fertilizer, and responses to P and S fertilizer.

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23.
Brix, H. and A.K. Mitchell. 1983. Thinning and nitrogen fertilization effects on sapwood development and relationships of foliage quantity to sapwood area and basal area in Douglas-fir. Canadian-Journal-of-Forest-Research 13(3): 384-389.

Keywords:      fertilization

                        thinning

                        tree morphology

                        tree physiology

Abstract: A 24-yr-old stand in British Columbia was treated in 1971-72 with various intensities and combinations of N fertilization and thinning. For 5-9 yr after treatments, trees were sampled to determine effects on foliage quantity and sapwood characteristics at varying stem ht. together with their relationships. Sapwood width remained relatively constant up the stem where heartwood was present, but the number of annual rings it contained decreased with ht. The sapwood width at b.h. increased with stem diam.; treatments had little effect on % sapwood at b.h. The ratio of foliage mass to sapwood cross-sectional area changed for different portions of the crown and was lower when based on sapwood area at b.h. than at base of live crown. Significant linear relationships of foliage mass and area to sapwood area at b.h. were found, but relationships of foliage to b.a. were just as close for all treatments; treatments significantly affected these relationships with control trees having the lowest regression slopes.

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24.
Brix, H. and A.K. Mitchell. 1986. Thinning and nitrogen fertilization effects on soil and tree water stress in a Douglas-fir stand. Canadian-Journal-of-Forest-Research 16(6): 1334-1338.

Keywords:      thinning

                        fertilization

                        soil properties

                        tree physiology

Abstract: Soil and tree water potentials were studied for 10 yr in a Douglas fir stand near Shawnigan Lake, British Columbia that was treated when 24 yr old with heavy thinning (removing superscript 2/3 of b.a.) and/or fertilization with 448 kg N/ha as urea. Control plots were not thinned or fertilized. Throughout the 10 yr, thinning increased soil water potential during the dry summer periods (July-early Oct.) by as much as 1 MPa. The effect of fertilization on soil water potential was slight and nonsignificant, and only apparent towards the end of the study in spite of large increases in leaf area (50% after 7 yr). Fertilization increased water use efficiency. The favourable soil water conditions produced by thinning led to improved shoot water potential only during predawn and early morning. Removal of understorey in a thinned and fertilized plot did not affect soil or shoot water potential.

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25.
Carr, W.W. 1987. Restoring productivity on degraded forest soils: two case studies. B.C. Ministry of Forests FRDA-Report 002. vi + 21 p.

Keywords:      site preparation

                        fertilization

                        tree physiology

                        growth

                        soil properties

Abstract: The use of green fallowing was studied at 2 sites, viz. (a) a coastal site at Koksilah, 15 km NW of Shawnigan Lake, British Columbia, where extensive subsoil exposure had resulted from roading operations in a highly productive Douglas fir (Pseudotsuga menziesii) and western hemlock (Tsuga heterophylla) stand, and (b) an inland site 30 km S. of Vanderhook including several landings and skid roads which had been deep-ripped to a depth of 50 cm, reducing soil density to 1350 kg/msuperscript 3. Plots at (a) were seeded in 1976 at 100 kg/ha with a grass/legume mixture including 3 spp. of Trifolium and Lotus corniculatus, and received NPK (10:30:10) at 450 kg/ha. Site nutrient levels improved substantially over 5 yr with N showing the greatest gains, and Douglas fir seedlings (1+2) planted in 1977 responded with increased foliar N and K contents and 300% greater ht. growth. Plots at (b) were seeded in 1981 at 40 kg/ha with a legume mixture of 3 spp. of Trifolium, Medicago sativa and L. corniculatus, and received NPK (19:19:19) at 300 kg/ha. Nutrient gains were found after 2 yr for P, K and especially N, and although foliar nutrient contents and growth of lodgepole pine (Pinus contorta) seedlings showed no increase, the enhancement of site nutrient capital is considered to be a gain likely to benefit commercial forestry production.

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26.
Carter, R. and K. Klinka. 1992. Use of ecological site classification in the prediction of forest productivity and response to fertilisation. South-African-Forestry-Journal (160): 19-23.

Keywords:      fertilization

growth
soil properties

Abstract: The results are presented of two related studies in southern coastal British Columbia. A total of 149 study plots in even-aged immature Douglas fir (Pseudotsuga menziesii) stands were categorized into 6 ecologically similar site associations (indicated best by climax plant communities) and into 15 soil moisture and soil nutrient combinations, for the prediction of site index. Fifty-one of these plots were used to predict basal area growth response to nitrogen fertilizer (225 kg N/ha as urea). Regression models indicated strong correlations between Douglas fir site index (m (height)/50 yr) and both indirect and direct measures of soil moisture and nutrient regimes (Rsuperscript 2 >0.72; p <0.01). Third-year basal area response to N fertilizer varied significantly among site associations (Rsuperscript 2 = 0.60; p <0.01). Site index also showed a significant relation with third-year basal area response (Rsuperscript 2 = 0.52; p <0.01), while the best predictive model included site associations and pretreatment foliar N and sulfate-S (Rsuperscript 2 = 0.64; p <0.01). It is suggested that the approach and methods of the study are applicable to predicting site-specific growth performance and response to fertilization of other tree species, including eucalypts.

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27.
Carter, R. and R. Scagel. 1989. The effects of stand density and fertilization on stand development in immature coastal Douglas-fir. B.C. Ministry of Forests FRDA-Report 094. i + 15 p.

Keywords:      fertilization

                        thinning

                        growth

                        tree morphology

Abstract: The effects of urea fertilizer were studied in Pseudotsuga menziesii stands spaced to 500, 750 and 1000 stems/ha in coastal British Columbia.

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28.
Carter, R.E. and R.P. Brockley. 1990. Boron deficiencies in British Columbia: diagnosis and treatment evaluation. Forest-Ecology-and-Management 37(1-3): 83-94.

Keywords:      fertilization

                        growth

                        tree/stand health

Abstract: Two case-studies are presented, outlining methods of diagnosis and treatment evaluation of boron deficiencies in a Douglas fir (Pseudotsuga menziesii) stand in coastal southern British Columbia and a lodgepole pine (Pinus contorta) stand near Burns Lake, in the interior of the province. Site conditions commonly associated with B deficiency are outlined, and relations between dormant-season foliar B concentration and growing-season precipitation and moisture stress are suggested. Diagnostic methods used in the study include examination of deficiency symptoms, and foliar-analysis techniques; evaluation of response for corrective fertilizer treatments is based on changes in frequency and severity of deficiency symptoms and growth responses, measured by graphical-analysis and changes in shoot length between treatments. Boron deficiencies and response to B fertilizers are difficult to confirm. Results of graphical-analysis and examination of frequency and severity of deficiency symptoms were inconclusive, while changes in shoot length identified a measurable response in the Burns Lake fertilizer trial with lodgepole pine. Deficiencies appear to be acute rather than chronic, and may not occur in untreated control trees for several years after establishment of fertilizer trials. Alternative causes for deficiency symptoms are also common, further complicating diagnosis and evaluation of response to treatment. It is concluded that all future trials should include nitrogen and/or other limiting nutrients with and without B to aid in identification of acute B deficiencies and deficiencies induced by increasing growth.

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29.
Carter, R.E., E.R.G. McWilliams and K. Klinka. 1998. Predicting response of coastal Douglas-fir to fertilizer treatments. Forest-Ecology-and-Management 107(1/3): 275-289.

Keywords:      fertilization

growth
soil properties

Abstract: A broadly-based, intensive Douglas-fir (Pseudotsuga menziesii) fertilizer experiment throughout southern coastal British Columbia (48 sites) was used to examine 3- and 6-yr crop tree growth responses to prescribed fertilizer applications (N alone, and N + additional nutrients identified as potentially deficient by foliar analysis). Absolute and relative basal area responses were evaluated in relation to site associations of the provincial ecosystem classification system, site index (SI), and a large number of site and stand chemical and physical properties. Few of the site and stand variables examined as possible response prediction criteria appeared to have any real utility. The strongest relationships found were between relative basal area response and (1) site index (Rsuperscript 2 0.46 for both 3- and 6-yr responses), (2) mineral soil mineralizable-N (Rsuperscript 2 0.50 and 0.46 for yr 3 and 6 responses, respectively), and (3) total mineralizable-N (Rsuperscript 2 0.47 and 0.50 for yr 3 and 6 responses, respectively). In all cases average relative response declined with increasing site quality. However, there were highly productive sites (SI50 <more or =>35 m) characterized by an absence of growing-season water deficits and relatively low foliar N concentrations (12-13 g/kg) which showed significant fertilizer responses. These sites are where the greatest financial returns from fertilizing may be realized. Relationships identified between site and stand variables and basal area responses were, in many cases, different from those found by other researchers for coastal Douglas fir. This brings the portability of identified relationships into question.

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30.
Chapman, R.J. 1984. Growth, nitrogen content and water relations of sludge-treated Douglas-fir seedlings. Forestry-Abstracts 45(7): 385-386.

Keywords:      fertilization
growth

                        tree physiology

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31.
Chappell, H.N., D.W. Cole, S.P. Gessel and R.B. Walker. 1991. Forest fertilization research and practice in the Pacific Northwest. Fertilizer-Research 27(1): 129-140.

Keywords:      fertilization

                        soil properties

Abstract: A review showed that most Pacific Northwest USA Douglas-fir (Pseudotsuga menziesii) forest sites are nitrogen deficient. Mineral cycling research has shown high C:N ratios and low nitrification rates for soils in the region. Research and development projects in the Pacific Northwest have produced an information base that is used to select sites and stands for fertilization and to forecast growth after treatment. Much of the basis for operational fertilization programmes in western Oregon and Washington comes from cooperative research; current activities for these programmes are directed toward improving site-specific response information. Forest fertilization in the Pacific Northwest has become a major silvicultural practice over the past two decades. Forest industry and government organizations managing forest lands in western Oregon and Washington apply nitrogen fertilizer to Douglas-fir stands over a range of soil and stand types. About 50 000 to 55 000 ha are fertilized each year, and future programmes will probably be of similar magnitude. Most current plans for management regimes including fertilization require multiple applications.

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32.
Chappell, H.N., C.E. Prescott and L. Vesterdal. 1999. Long-term effects of nitrogen fertilization on nitrogen availability in coastal Douglas-fir forest floors. Soil-Science-Society-of-America-Journal 63(5): 1448-1454.

Keywords:      fertilization

                        soil properties

Abstract: The aim of this study was to determine if N availability was elevated 8 to 12 years after repeated N fertilization, and if the effects of N fertilization were related to the soil N capital. Rates of N cycling in control and fertilized plots of Douglas-fir (Pseudotsuga menziesii) in Oregon, USA, were compared by measuring net N mineralization rates in forest floors and by estimating rates of N turnover from the litterfall/forest floor ratio. Litterfall N contents, litter N concentrations, and rates of N turnover increased along the gradient in soil N capital in both control and fertilized stands. Fertilization did not affect litterfall N content, but C:N ratios of litter and forest floors were significantly lower in fertilized stands along the gradient. Turnover rates of N in the forest floors were not higher in fertilized plots than in control plots, nor were rates of net N mineralization affected by fertilization. Net nitrification rates were higher in some of the plots that received 1120 kg N ha-1 than in control plots. Nitrogen fertilization did not result in a sustained increase in N cycling and N availability analogous to a higher site N capital, and the effect of N fertilization was not related to the initial soil N capital of these sites.

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33.
Cochran, P.H., W. Lopushinsky and P.D. McColley. 1986. Effect of operational fertilization on foliar nutrient content and growth of young Douglas-fir and Pacific silver fir. Pacific-Northwest-Research-Station,-USDA-Forest-Service Research-Note PNW-RN-445. 10 p.

Keywords:      fertilization

                        tree physiology

                        growth

Abstract: During 1979-80, sulfated urea (pelletted) was applied to conifer stands in the Wenatchee National Forest, west of Cle Elum, Washington. Nitrogen concn. in current needles of Pacific silver fir (Abies amabilis) showed a significant 1.9-fold increase after fertilizer treatment compared with a non-significant 1.3-fold increase in Douglas fir (Pseudotsuga menziesii). A significant 2.5-fold increase in foliar N also occurred in bracken (Pteridium aquilinum). Analysis of foliage from untreated trees indicated N deficiency in A. amabilis, but N concn. in Douglas fir was above threshold values. Fertilizer treatment did not affect foliar S in either species, but increased needle surface area for A. amabilis. Tree diam. growth, stand b.a. growth and vol. growth were all increased by fertilizer treatment.

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34.
Cole, D.W., M.L. Rinehart, D.G. Briggs, C.L. Henry and F. Mecifi. 1984. Response of Douglas fir to sludge application: volume growth and specific gravity. In Proceedings of the Technical Association of the Pulp and Paper Industry 1984 Research and Development Conference, Appleton, Wisconsin, September 30-October 3. pp. 77-84.

Keywords:      fertilization

                        thinning

                        growth

                        wood quality

Abstract: In 1977 and 1980 municipal sludge was applied to a 60-yr-old lowland Douglas fir stand in Washington State. Application procedures and rates and suitable sites for treatment are described. There was a 6 yr av. diam. growth response of 93% in unthinned and 48% in thinned stands treated with 142 t/ha sludge and a vol. growth response of 53 and 42%, respectively. The accelerated rate of growth has not shown signs of decreasing since treatment. Relative density of sludge-grown wood was 10-15% less than that of untreated wood, but within the range for Douglas fir grown on higher land. This is thought to be a result of the change in forest site produced by the sludge treatment.

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35.
Colinas, C., R. Molina, J. Trappe and D. Perry. 1994a. Ectomycorrhizas and rhizosphere microorganisms of seedlings of Pseudotsuga menziesii (Mirb.) Franco planted on a degraded site and inoculated with forest soils pretreated with selective biocides. New-Phytologist 127(3): 529-537.

Keywords:      planting operations

                        fertilization

                        soil properties

                        mycorrhizal response

Abstract: Inoculation of planting holes with small amounts of soil from a mature forest or a plantation can improve formation of ectomycorrhizas on Pseudotsuga menziesii seedlings in degraded clearcuts in southwestern Oregon. To determine the component(s) of transferred soil responsible for increased ectomycorrhiza formation, soil from a clearcut, a mature forest and a plantation was treated with one of the following: (1) fertilizer to test for the effect of nutrients, (2) dimethoate and carbofuran to test for the effect on microarthropods or nematodes, (3) fumagillin to test for the effect on protozoa, (4) captan to test for the effect on fungi, (5) penicillin and oxytetracycline to test for the effect on bacteria, (6) pasteurization to test for the effect of active forms of organisms, (7) Tyndallization to test for the effect of resting forms of organisms, or (8) water as a control. The effect was studied of inoculation with soil subjected to these treatments on number and types of ectomycorrhizas, on length of active mycelium, and on number of active bacteria in the rhizosphere. Inoculation with untreated forest or plantation soils increased the number of ectomycorrhizas but did not change the mycorrhizal types present. Most agents had different effects in different soils. Inoculation with pasteurized and Tyndallized clearcut and plantation soils increased the number of Rhizopogon- and Thelephora-type ectomycorrhizas and decreased the number of active bacteria, as did untreated forest soil. It is hypothesized that the role of the soil transfer is to provide a rhizosphere environment free from a deleterious organism present in the clearcut. In this environment, beneficial organisms present in the clearcut or brought in with the seedling from the nursery can proliferate.

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36.
Colinas, C., D. Perry, R. Molina and M. Amaranthus. 1994b. Survival and growth of Pseudotsuga menziesii seedlings inoculated with biocide-treated soils at planting in a degraded clearcut. Canadian-Journal-of-Forest-Research 24(8): 1741-1749.

Keywords:      planting operations

                        fertilization

                        growth

                        tree/stand health

                        tree morphology

Abstract: To determine the factors of transfer soils responsible for increased seedling survival and growth, planting holes, at a site in SW Oregon, were inoculated with forest, plantation, and clear-cut soils subjected to one of 8 treatments: (i) treated with fertilizer to test for effects of nutrients; treated with biocides to test for effects of (ii) grazers (microarthropods or nematodes), (iii) protozoa, (iv) fungi, or (v) bacteria; (vi) pasteurized; (vii) Tyndallized; or (viii) untreated. Douglas fir (Pseudotsuga menziesii) seedlings were planted in June 1990 and seedling growth and survival was assessed in December 1990. Survival was increased by inoculation with untreated plantation soils, but not if they were fertilized or treated with dimethoate + carbofuran (grazercide), fumagillin (protozoacide), or oxytetracycline + penicillin (bactericide). Addition of untreated forest soil did not increase survival. For all soils, survival was increased by captan (fungicide), pasteurization and Tyndallization. Untreated plantation and forest soil transfers increased dry weights whereas neither did when treated with dimethoate + carbofuran. Dry weights of seedlings given clear-cut soil were increased by fertilization, pasteurization and Tyndallization of the soil; the latter two treatments also increased the number of short roots. It is hypothesized that stimulation of seedling growth by soil transfers was related to an increased rate of nutrient mineralization due to microbivorous soil animals contained within the transfer soils. Soil transfers may have enhanced seedling survival by at least two mechanisms: (i) by providing a safe site for beneficial rhizosphere organisms to proliferate, free from competing organisms that have proliferated in the clear-cut soil; and (ii) through volatile organic compounds, especially ethylene, that stimulated seedling root growth.

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37.
Crouch, G.L. and M.A. Radwan. 1981. Effects of nitrogen and phosphorus fertilizers on deer browsing and growth of young Douglas-fir. Pacific-Northwest-Forest-and-Range-Experiment-Station,-USDA-Forest-Service. Research-Note PNW-RN-368. 15 p.

Keywords:      fertilization

                        tree/stand health

growth
tree physiology

Abstract: N and P fertilizers were applied in March 1968 singly or in combination at a rate equivalent to 200 lb/acre of N or P to young trees (2-5 ft tall) in Oregon and Washington. Trees were examined and measured for up to 4 yr. In the first year after treatment trees given the N-only fertilizer in Washington were more heavily browsed by black tailed deer (Odocoileus hemionus columbianus), but this effect disappeared in the second year. Height growth was increased by N-only treatment in taller trees in Oregon after 1 and 2 yr, but the effect had disappeared after 4 yr. Total N content was significantly increased by the N and N + P treatments in the first year, but this effect also disappeared after 2 yr. The amounts of moisture, ash, Ca and P, and diam. growth were not affected by any treatments.

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38.
Curtis, R.O., G.W. Clendenen and D.J. DeMars. 1981. A new stand simulator for coast Douglas-fir: DFSIM user's guide. Pacific-Northwest-Forest-and-Range-Experiment-Station,-USDA-Forest-Service General-Technical-Report PNW-GTR-128. ii + 79 p.

Keywords:      planting operations

                        thinning

                        precommercial thinning

                        commercial thinning

                        fertilization

                        yield

                        computer modeling

Abstract: A description of a computer program, written in FORTRAN IV, for simulating managed stands. The program has been developed from remeasured plot data contributed by many organizations in the Pacific Northwest USA. It can produce yield tables which include estimates of effects of initial spacing, precommercial and commercial thinning and addition of N fertilizer. Topics discussed include program limitation and potential for further development. Appendices include operating instructions and notes on testing. The program is available from the authors on request.

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39.
Dangerfield, J. and H. Brix. 1981. Comparative effects of ammonium